Clada (Clada) njakai, Zahradnik, Petr & ryzna, Milos, 2018

Zahradnik, Petr & ryzna, Milos, 2018, Nine new species of Clada from Madagascar (Coleoptera, Ptinidae), ZooKeys 806, pp. 121-140 : 121

Clada (Clada) njakai sp. n. Figs 7, 16, 25, 34

Type material.

Holotype male: Madagascar, Toliara prov., Tsimanampetsotsa N. P., Mitoho camp, 24°02.898'S, 43°45.138'E, 10 m a. s. l., 12.-13.i.2014, M. Trýzna leg. Paratypes (15): 1 male: the same data as holotype; 13 males: Madagascar, Toliara prov., Tsimanampetsotsa N. P., Andranovao camp, 24°01.505'S, 43°44.306'E, 15 m a. s. l., 14.-15.i.2014, M. Trýzna leg. (FGMRI 7 ex., LBVC 2 ex., MTDC 2 ex., NHMUK 2 ex.).

Differential diagnosis.

This species is similar to C. (C.) humeralis Pic, 1926, but differs by the lighter colour of the elytra and missing lighter humeri. Fully differs by shape of the aedeagus. For differences from other Madagascan species, see key.

Description.

Male (holotype). Elongate-elliptical, transversally convex. Body length 5.3 mm, maximum width 2.1 mm (Figure 7). Ratio length:width of elytra 1.8. Whole body dark brown, only antennae, maxillary and labial palpi and humeri on elytra moderately lighter. Pubescence yellowish-white.

Head matt-shiny, with double punctation - one coarse, dense, umbilicate, distance between punctures approximately the same as their diameter; other one very fine, punctures almost touching. Pubescence recumbent or semi-erect, short, inclined backwards; on sides of head semi-erect and long, inclined forwards. Clypeus with shallow transverse depression. Eyes large, globular with short erect sparse pubescence. Frons twice as wide as diameters of eye, from dorsal view. Antennae consisting of eleven antennomeres, 4th to 10th pectinate (Figure 25). 1stantennomere robust, twice as long as wide; 2nd smallest, only one-half length of 1st, as long as wide, slightly narrower than 1st. 3rd serrate, as long as wide; 4th and 5th 1.8 times wider than long; 6th and 7th 1.5 times wider than long; the 8th and 9th 1.3 times wider as long; and 10th 1.1 times as wide as long. Apical antennomere is longest, oblong oval, 4.3 times as long as wide. All antennomeres on margin with short erect dense setae. Apical maxillary palpomere long, spindle shaped.

Pronotum convex, matt-shiny, rounded, transverse (ratio length:width of pronotum 0.7); widest at 2/3 posteriorly. Base of pronotum finely bordered. Middle of pronotum at base with blunt small swlling, posteriorly slightly sharpened. Surface of pronotum with double punctation, one coarse, dense, umbilicate, distance between punctures approximately the same as their diameter; other one is very fine, punctures almost touching. Pubescence long, sparse, recumbent, inclined more or less to middle of pronotum.

Scutellum triangular, narrow, 1.3 times as long as wide, very densely recumbent pubescence, inclined backwards, surface is not visible.

Elytra oval, transversally convex, shining, with distinct humeri. Each elytron with five fine costae, almost invisible, but apex more distinct. Surface of elytra with double punctation, one coarse, dense, umbilicate, distance between punctures approximately the same as their diameter; other one is very fine, punctures almost touching. Pubescence relatively sparse, recumbent or semi-erect, inclined backwards. Posterior margin of each elytron with approximately 25 small teeth.

Legs stout, with short and dense recumbent pubescence. All tarsi robust, same length as tibia. 1st metatarsomere as long as 2nd and 3rd together, same width, slightly emarginate on top, 4th slightly shorter than previous, more emarginate, almost to 2/3 of their length. 5th is same length as 3rd and 4th together, rectangular, wider on the top, with two large claws, without teeth.

For aedeagus see Figure 34.

Female. Unknown.

Variability.

Body length 4.7-7.1 mm, maximum width 1.8-2.8 mm.

Name derivation.

Patronym, dedicated to Adolphe Randrianjaka (University of Antananarivo, Faculty of Sciences, Department of Entomology), whom we called Njaka.

Biology.

Unknown. All specimens were collected at light.

Distribution.

This species is found in the southwestern part of Madagascar (Figure 16).