Clidicus mawarensis, Jałoszyński, 2020

Jałoszyński, Paweł, 2020, A new species of Clidicus in East Malaysia (Coleoptera, Staphylinidae Scydmaeninae), Zootaxa 4790 (1), pp. 187-192 : 189-191

Clidicus mawarensis sp. n.

( Figs 7–12 )

Material studied. HOLOTYPE: ♂: MALAYSIA (SABAH), with two labels: “MALAYSIA-Sabah / Mawar wa- terfall / near Patau village / 305.-31.5.98, P.Hlaváč ” [white, printed], “ CLIDICUS / mawarensis m. / P. Jałoszyński, 2020 / HOLOTYPUS ” [red, printed] ( NMPC).

Diagnosis. Body large, BL> 6.5 mm; punctures on pronotum separated by distances subequal to their diameters; most flagellomeres elongate; aedeagus stout and massive, with short, slender and slightly asymmetrical parameres (one slightly longer than the other), with broad apex of median lobe and complex apical structures, with a pair of lateral apical hook-like projections at sides of gonopore-bearing median projection and with ventral subtrapezoidal shield-like plate with deeply emarginate apex.

Description. Body of male ( Fig. 1 ) elongate and slender, head flattened, prothorax and elytra strongly convex; legs and antennae long and slender, BL 6.75 mm; cuticle glossy, pigmentation uniformly reddish brown, setae indistinctly lighter than cuticle.

Head ( Fig. 2 ) broadest distinctly behind middle of tempora, HL 1.13 mm, HW 1.50 mm; tempora in dorsal view nearly 4 times as long as eyes, nearly straight in slightly more than anterior half and weakly divergent posterad, posteriorly strongly and evenly rounded; vertex with a deep subtrapezoidal posteromedian impression and a pair of indistinct posterior pits marking sites where dorsal tentorial arms are fused with dorsal wall of head. Sides of vertex convex, frons near each antennal insertion slightly impressed, entire frons and vertex with a narrow median longitudinal groove, supraantennal tubercles small but distinctly elevated; clypeus transverse and convex, demarcated from frons by a deep and distinct frontoclypeal groove; compound eyes small, nearly round, strongly convex but in dorsal view not projecting from head silhouette, finely faceted. Punctures on vertex and frons sparse and fine, inconspicuous; supraantennal tubercles virtually impunctate. Setae moderately long, sparse, erect. Antennae slender, scape slightly longer than head, antennomeres II–VIII elongate (VIII indistinctly so), IX–X as long as broad, XI indistinctly transverse; AnL 3.63 mm.

Labrum with shallow subtriangular anteromedian notch.

Pronotum with subglobose disc and short posterior ‘collar’, broadest near anterior third; PL 1.63 mm, PW 1.55 mm; ‘collar’ demarcated by a narrow transverse groove with twelve pits increasing in diameter toward sides. Punc- tures on disc denser than those on head, in median area separated by spaces subequal to diameters of punctures; setae moderately long, dense, and suberect. Cuticle between punctures on pronotal disc as glossy as that on head.

Elytra together oval and strongly elongate, broadest slightly in front of middle, EL 4.00 mm, EW 2.15 mm, EI 1.86; each elytron with five dorsal and one lateral rows of large and deep, densely distributed pits, cuticle between pits glossy, between rows with fine and sparse setiferous punctures; setae longer and slightly thicker than those on head and pronotum, moderately dense, suberect. Apex of each elytron obliquely truncated, with lateral corner form- ing short denticle; adsutural corner weakly projecting posterad.

Hind wings present, longer than elytra but narrow and apparently not functional.

Legs long and slender, unmodified.

Abdominal sternite IX ( Fig. 3 ) with broadly V-shaped subapical transverse carina, area behind carina densely setose.

Aedeagus ( Figs 4–5 ) relatively stout, in lateral view weakly curved, AeL 1.33 mm; parameres short, slightly asymmetrical, one slightly longer than the other, their apical portions projecting beyond apex of median lobe nearly parallel; flagellum differentiated from ductus ejaculatorius by a rapid and strong broadening of walls near distal fourth of median lobe. Apex of median lobe (between parameres; Figs 7–12 ) broad and with median rounded projection with broad gonopore opening ventrally, flanked by a pair of robust tooth-like projections directed ventrad, ventral apical region of median lobe with shield-like plate with deeply emarginate distal margin. Sperm pump ( Fig. 6 ) 0.80 mm in length, with two large ‘funnels’, distal funnel much shorter than proximal one.

Female. Unknown.

Distribution. Northern Borneo.

Etymology. After Mawar waterfall.

Remarks. Clidicus mawarensis has large-bodied adults, which are externally similar to several congeners inhabiting SE Asia. Examination of the aedeagus is necessary to identify the new species. The male copulatory organ of C. mawarensis is most similar to that of C. ganglbaueri Reitter, 1887 , known to occur in Indonesia (Java and Sumatra). In both species the median lobe is relatively massive and stout, the parameres are slightly asymmetrical, and the sperm pump has a similar shape. However, in C. mawarensis the entire aedeagus is much slenderer in lateral view; the median lobe is broadest in the subapical region (in sub-basal region in C. ganglbaueri ); the subapical region bears a small shield-like ventral plate with pointed distolateral corners (in C. ganglbaueri an elongated ventral plate with rounded distolateral corners); and the parameres are distinctly shorter in relation to the median lobe and constricted (long and gradually narrowing in C. ganglbaueri ). Moreover, the elytra of males are more elongate in C. ganglbaueri .

Figures

FIGURES 7–12. Clidicus mawarensis sp. n. apical portion of median lobe in dorsal (7–8), ventral (9–10) and lateral (11–12) views. Abbreviations: cp, copulatory piece; fl, flagellum; gon, gonopore; hlp, hook-like projection.

FIGURES 1–3. Clidicus mawarensis sp. n. Habitus of male (1); head of male in dorsal view (2), apex of male terminalia in ventral view (3). Abbreviation: st9, sternite IX.

FIGURES 4–6. Clidicus mawarensis sp. n. Aedeagus in dorsal (4) and lateral (5) views; sperm pump (6).