Coccidencyrtus solivus, Noyes, 2023

Noyes, John Stuart, 2023, ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera), Taxonomic Monographs on Neotropical Hymenoptera (Oxford, England) 2 (11), pp. 1-921 : 120-121

Coccidencyrtus solivus sp.nov.

( Figs 275-280; Hab. E 46)

DIAGNOSIS. Female (overall length about 0.8mm): scape mostly dark brown; pedicel and flagellum pale yellow; all coxae white; fore femur and tibia white to very pale yellow; mid femur white with a subapical brown band; mid tibia white to pale orange with a narrow subbasal brown band; hind femur white with a brown subapical band; hind tibia white with a narrow subbasal brown band; gaster dorsally dark brown, medially and ventrally pale orange, hypopygium brown; frontovertex slightly less than half head width; antenna ( Fig. 275) with scape about 2.5X as long as broad, funicle with F1 narrower than pedicel, F1-F6 relatively small, F6 largest and subquadrate; linear sensilla present on F4-F6 or missing from F5; clava very slightly longer than funicle, about 3X as long as broad; palp formula 3-2; scutellum ( Fig. 279) with 3 pairs of setae and longitudinally elongate polygonally reticulate to striate-reticulate sculpture, deeper than that on mesoscutum; mid basitarsus with 3 or 4 ventral pegs; mid tibial spur about as long as basitarsus; fore wing with stigmal vein slightly longer than postmarginal vein but shorter than marginal vein ( Fig. 276); hypopygium reaching about 0.7X towards apex of gaster; ovipositor about 4X as long as gonostylus and slightly exserted with exserted part slightly shorter than mid tibial spur. Male: unknown.

Female (holotype): length, including ovipositor, 0.79mm; excluding ovipositor, 0.73mm ( CPD) .

Head dark brown to black with a weak to moderate metallic lustre; occiput with a weak brassy sheen; frontovertex generally metallic dark green mixed coppery, area between posterior ocelli and occipital margin coppery; area above scrobes coppery; scrobal area, temple and gena with a weak coppery sheen, interantennal prominence coppery purple; frontovertex with a few, inconspicuous, scattered, brown setae; gena and interantennal prominence with a few scattered, very inconspicuous, brown setae; maxillary palpus white; radicle orange-brown to dark brown; scape mostly dark brown, apical 0.3X pale yellow; pedicel and flagellum pale yellow, clava slightly dusky; thorax dark brown to black with a metallic lustre; pronotum with a weak coppery sheen; mesoscutum relatively bright metallic dark green mixed coppery especially near margins and in posterior half; tegula dark purple-brown with a weak brassy sheen; axilla with a weak coppery purple and brassy sheen; scutellum dorsally virtually matt, but with a very weak coppery purple sheen, extreme side and apex weakly shining and slightly brassy; mesopleuron with a moderate purple sheen; all coxae white, hind coxa slightly dusky proximally; fore femur and tibia white to very pale yellow, tarsus slightly dusky, pale orange; mid femur white with a narrow, incomplete, dark brown subapical band, tibia white to pale orange with a narrow, subbasal dark, brown band, spur and tarsus very pale orange, apical tarsomere slightly dusky; hind femur white with a slightly oblique, narrow, brown, subapical band, tibia white with a narrow, brown, subbasal band, tarsus very pale yellow, apical tarsomere slightly dusky; wings mainly hyaline, but fore wing with a small, very faint, pale fuscous patch below the marginal and stigmal veins( Fig. 276); propodeum dark brown with a weak purple sheen; gaster dorsally purple-brown with coppery purple and green reflections, Gt1-Gt3 medially pale yellow, side and venter mostly dark brown, but basal sternites pale yellow; gonostylus dark brown.

Head ( Fig. 277) about 2.3X as wide as frontovertex, in profile about 1.6X as high as deep, tangent to frontovertex forming an angle of about 105° with that of face; occipital margin more or less sharp, not carinate; no distinct oval depression adjacent to eye near occipital margin; ocelli forming an angle of about 95°; frontovertex not widening in front of anterior ocellus, with distinct polygonally reticulate sculpture of mesh size subequal to, or a little smaller, than eye facet, not shallower above top of scrobes; temple and gena with slightly shallower, longitudinally elongate, fine polygonally reticulate sculpture; top of scrobes with distinct, regular, polygonally reticulate sculpture; interantennal prominence and mouth margin with slightly shallower, polygonally reticulate sculpture; scrobes shallow, indistinctly margined, broadly ∩-shaped; interantennal prominence dorsally rounded; antenna as in Fig. 275; scape very slightly broadened and flattened, about 2.7X as long as broad; funicle with F1-F5, relatively small, F6 largest and quadrate or very slightly longer than broad; linear sensilla present on F4-F6 or missing from F5; clava moderately slender, about 3.2X as long as broad, about 1.5X as wide as F6, not downcurved ventrally, very slightly longer than funicle, sutures parallel, sensory area relatively small, at apex only; eye slightly overreaching occipital margin, clothed with dense, conspicuous setae, each slightly longer than diameter of facet; malar sulcus absent; clypeal margin evenly quite strongly concave; mandible with two acute teeth and an upper obtuse tooth; palp formula 3-2. Relative measurements: HW 45, HH 34.5, FV 20, POL 10, OOL 2.5, OCL 1, AOL 7, EL 27, EW 22.5, MS 12.5, SL 16, SW 6.

Thorax ( Fig. 279) with pronotum hardly exposed behind head, its posterior margin broadly concave, not angular medially; pronotum and mesoscutum with polygonally reticulate sculpture that is about as deep as that in ocellar area, and of similar mesh; axilla with similar sculpture to mesoscutum but clearly shallower; scutellum dorsally with very fine, slightly longitudinally elongate, polygonally reticulate sculpture that is a little deeper than sculpture on mesoscutum, extreme side and apex completely smooth and shiny; visible part of mesoscutum about 1.8X as broad as long; scutellum about 1.3X as broad as long, apex with a pair of slender, elongate setae; hind femur about 3.5X as long as broad; fore wing with venation and setation as in Figs 276, 278; propodeum ( Fig. 279) medially about 0.05X as long as scutellum. Relative measurements: FWL 101.5, FWW 41; HWL 69, HWW 14.5.

Gaster with hypopygium reaching about 0.7X to apex; last tergite about 0.7X as long as mid tibia, with apex broadly rounded; hypopygium Fig 280; ovipositor exserted, the exserted part about 0.8X as long as mid tibial spur or about 0.25X mid tibia. Relative measurements: OL 86, GL 22 [MT 75].

Variation. Only holotype examined.

Male. Unknown.

DISTRIBUTION. Costa Rica.

HOSTS. Unknown.

MATERIAL EXAMINED.

Type material. Holotype E: COSTA RICA, Guanacaste, Guanacaste NP (error for Santa Rosa NP ), near HQ, MT/YPT, 2-10.iii.1990 (J.S. Noyes) . Holotype in NHMUK .