Coniopteryx (Coniopteryx) evellana, Sziraki, 2015

Sziráki, György, 2015, Identity Of Coniopteryx Madagascariensis Meinander, 1974 (Neuroptera: Coniopterygidae), With Description Of Three New Species, Acta Zoologica Academiae Scientiarum Hungaricae 61 (2), pp. 135-146 : 139-142

Coniopteryx (Coniopteryx) evellana sp. n.

( Figs 8–13)

Coniopteryx (C.) madagascariensis sensu MeInandeR 1998, Figs 17B, 17E, 17G, 17I, 17K.

Examined material – Holotype: male, Madagascar, Fianarantsoa Province, Ranomafana National Park, Belle Vue at Talatakely, 21°15’59” S, 47°25’13” E, 1020 m a.s.l., secondary tropical forest, 16. X – 8. XI. 2002, Malaise trap, leg.: HH; deposited in the collection of CAS. Paratypes : 3 males, Madagascar, Antanarivo Province, 3 km NE of Andranomay , 18°28’24” S, 47°57’36” E, 1300 m. a.s.l., mountain rainforest, 5-13. XII. 2000 , Malaise trap, leg.: F, G et al.; 1 male, Madagascar, Antsiranana Province, Marojejy National Park, Manatenia River , 8.2 km NNW of Manantenia, 14°26’12” S, 49°46’30” E, 450 m a.s.l., rain forest, 12-25. XI. 2003 , Malaise trap, leg.: F et al.; 1 male, Madagascar, Diego – Suarez Province, Montagne d’Ambre National Park , 12°31’ S, 49°11’ E, 975 m a.s.l., 21-25. I. 2001 , Malaise trap, leg.: I, ES et HH; 1 male, Madagascar, Fianarantsoa Province, Forêt d’ Ambalagoavy nord Ikongo, Ambatombe, 21°49’39” S, 47°20’20” E, 625 m . a.s.l., (day?) XI. 2000, Malaise trap, leg.: HH et I; 7 males, Madagascar, Fianarantsoa Province, Ranomafana National Parc, Belle Vue at Talatakely , 21°15’59” S, 47°25’13” E, 1020 m. a.s.l., 8-15. XI. 2001 , secondary tropical forest, Malaise trap, leg.: HH; 13 males, same data but 15-22. XI. 2001 ; 16 males, same data but 22-28. XI. 2001 ; 6 males, same data but 28. XI – 6. XII. 2001 ; 3 males, same data but 6-15. XII. 2001 ; 1 male, same data but 15-21. XII. 2001 ; 4 males, same data but 21-24. XII. 2001 ; 11 males, same data but 24. XII. 2001 – 2. I. 2002 ; 1 male, same data but 2-10. I. 2002 ; 2 males, same data but 10-14. I. 2002 ; 3 males, same data but 14-21. I. 2002 ; 13 males, same data but 21-28. I. 2002 ; 1 male, same data but 28. I. – 4. II. 2002 ; 2 males, same data but 12-19. II. 2002 ; 3 males, 19-26. III. 2002; 15 males, same data but 31. III – 7. IV. 2002 ; 1 male, same data but 7-14. IV. 2002 ; 7 males, same data but 14-23. IV. 2002 ; 1 male, same data but 28. IV – 5. V. 2002; 3 males, same data but 13-23. V. 2002; 2 males, same data but 23. V – 3. VI. 2002; 1 male, same data but 13-23. VI. 2002; 1 male, same data but 23. VI – 4. VII. 2002; 6 males, same data but 4-14. VII. 2002 ; 2 males, same data but 24. VII – 4. VIII. 2002 ; 14 males, same data but 16. X – 8. XI. 2002 ; 3 males, Madagascar, Fianarantsoa Province, Ranomafana National Parc, Jirama Water Works, 21°14’55” S, 47°27’08” E, 690 m . a.s.l., 22-28. XI. 2002, open area near a stream, Malaise trap, leg.: HH; 1 male, same data but 28. XI – 6. XII. 2001 ; 5 males, same data but 21-24. XII. 2001 ; 1 male, same data but 2-10. I. 2002 ; 3 males, same data but 10-14. I. 2002 ; 8 males, same data but 21-28. I. 2002 ; 1 male, same data but 16. X – 8. XI. 2002 ; 7 males, Madagascar, Fianarantsoa Province, Ranomafana National Park , radio tower, 21°15’30” S, 47°24’26” E, 1130 m. a.s.l., 16. X – 8. XI. 2001 , mixed tropical forest, Malaise trap, leg.: HH; 4 males, same data but 15-22. XI. 2001 ; 6 males, same data but 22-28. XI. 2001 ; 13 males, same data but 28. XI – 6. XII. 2001 ; 2 males, same data but 6-15. XII. 2001 ; 8 males, same data but 15-21. XII. 2001 ; 1 male, same data but 10-14. I. 2002 ; 15 males, same data but 14-21. I. 2002 ; 5 males, same data but 21-28. I. 2002 ; 1 male, same data but 12-19. II. 2002 ; 2 males, same data but 19-26. III. 2002 ; 2 males, same data but 26-31. III. 2002 ; 3 males, same data but 16-23. IV. 2002 ; 1 male, same data but 23-30. IV. 2002 ; 2 males, same data but 24. V – 4. VI. 2002; 5 males, same data but 14-24. VI. 2002; 4 males, same data but 15-25. VII. 2002 ; 1 male, same data but 18-27. II. 2003 ; 9 males, Madagascar, Fianarantsoa Province, Ranomafana National Parc, Vohiparara , 21°13’34” S, 47°22’11” E, 1110 m. a.s.l., 22-28. XI. 2001 , mountain rainforest, Malaise trap, leg.: HH; 36 males, same data but 28. XI – 6. XII. 2001 ; 2 males, same data but 6-15. XII. 2001 ; 2 males, same data but 15-21. XII. 2001 ; 4 males, same data but 24. XII. 2001 – 2. I. 2002 ; 1 male, same data but 10-14. I. 2002 ; 6 males, same data but 14-21. I. 2002 ; 3 males, same data but 21-28. I. 2002 ; 33 males, same data but 28. I – 4. II. 2002 ; 1 male, same data but 4-12. II. 2002 ; 1 male, same data but 12-19. II. 2002 ; 1 male, same data but 21-28. II. 2002 ; 1 male, same data but 12-19. III. 2002 ; 1 male, same data but 15-22. IV. 2002 ; 6 males, same data but 22-29. IV. 2002 ; 3 males, same data but 15-25. V. 2002; 3 males, 4-14. VI. 2002; 2 males, same data but 26. VI – 5. VII. 2002; 2 males, same data but 5-15. VII. 2002 ; 7 males, same data but 15-25. VII. 2002 ; 9 males, same data but 25. VII – 3. VIII. 2002 . Majority of paratypes are deposited in the collection of California Academy of Sciences , while 6 paratype specimens are housed in the Hungarian Natural History Museum, Budapest.

Diagnosis: Small bodied coniopterygid, with flattened projection between the antennae. First flagellar segment with an outstanding setose outgrowth. Processus terminalis truncated obliquely in ventral view. Median incision consists of a very shallow „v” shaped part and of a narrow engraving in the middle of it.

Description: Length of the body 1.3–1.8 mm. Head capsule medium brown, palpi light brown. Between the antennae the anterior edge of vertex has a prominent, laterally flattened projection. Eyes moderately large, black. Antennae dark brown, 0.9–1.2 mm, 26–29 segmented. Scape slightly shorter than wide, pedicel about as long as wide, flagellar seg- ments 2.4–3 times as wide as long. Pedicel has a very small, while the first flagellar segment has an outstanding outgrowth apically, with rather short setae ( Fig. 8). Ordinary hairs are arranged in two rather irregular rings on flagellar segments. Scale-like hairs on the whole surface of the pedicel, and in a very dense whorl on the apical half of the flagellar segments.

Legs, wings and the large part of thorax medium brown, sutures, apodemes and the shoulder spots dark brown. Length of fore wing 1.4–2.2 mm, of hind wing 1.4–1.7 mm. Pregenital part of abdomen light brown.

Male terminalia ( Figs 9–13) dark brown, well sclerotized. Hypandrium as long as high in lateral view. Processus terminalis distinct but truncated obliquely in ventral view. ventral lobe of the inner branch of stylus. Scales: 0.04 mm.

Therefore, medianincisionconsistsofaveryshallow „v” shapedpartandofanarrow engravinginthemiddleofit. Processuslateralissmallanddirectedcaudally. Itssetaeare rathershort. Anteriorapodemeofhypandriumalmoststraightinventralview, whilein lateralviewcurvedbackwards (andstonglywidened) dorsally. Gonarcuswellsclerotized, withabroadbasalpart. Stylusforked. Itsouterbranchnarrowinlateral, broadincaudal view, withaveryfinelyserratedending. Innerbranchofstyluswidewithapeculiarbroad, plate-like, subtriangularventrallobe. Theventralbridgebetweenthetwoinnerbranches moderatelysclerotized. Parameremoderatelylong; itscaudalpartturnedupwardssubapi- callyinanobtuseangle. Processusventralissmall, butdistinct. Processusapicalisrather smallalso, directedupwards, withatooth-likeupperandaplate-likelowerpart. Thepenis scleriteconsistsoftwomoderatelychitinized, bentrods.

Remarks: Coniopteryx (C.) evellana belongs to Coniopteryx lobifrons group sensuSZIRÁKI (2005), and – regardingtheprocessusterminalis, medianinci- sionanddistalpartofparamere – itresemblesto Coniopteryx (C.) oroszi Szirá- ki, 2004 fromThailand.

Themaindistinctivefeaturesofthenewspeciesare:

− pedicelofantennawithoutaverystrongseta (whichischaracteristicin C. oroszi );

− presenceoftheplate-likesubtriangularlobeoftheinnerbranchofthe stylus;

− theinnerbranchofthestyluswide;

− thesmallprocessuslateralisofhypandriumdirectedcaudally.

Etymology: Thenewspeciesisnamedafterthestructureofmedianinci-

sionofhypandrium.