Coprophilus (Zonyptilus) bimaculatus Luze, 1904

Gildenkov, Mikhail Yu., 2017, New synonymies of Coprophilus (Zonyptilus) pentatoma Fauvel, 1897 (Coleoptera: Staphylinidae: Oxytelinae), Zootaxa 4306 (1), pp. 130-136 : 133-134

Coprophilus (Zonyptilus) bimaculatus Luze, 1904 , syn. n.

( Figs. 5 , 12)

Coprophilus bimaculatus Luze, 1904: 79

Coprophilus bimaculatus: Bernhauer, 1908: 330 (as synonym of longicornis ) Coprophilus bimaculatus: Bernhauer and Schubert, 1911: 89 (as synonym of longicornis ) Coprophilus (Zonoptilus) bimaculatus: Tóth, 1991: 91 (lectotype designation, valid species) Coprophilus (Zonoptilus) bimaculatus: Tóth, 1992: 376

Coprophilus bimaculatus: Herman, 2001: 1312

Coprophilus (Zonyptilus) bimaculatus: Smetana, 2004: 511

Coprophilus (Zonyptilus) bimaculatus: Schülke and Smetana, 2015: 766

Type material examined. Lectotype ♂, Tadjikistan “TRKST. JAGNOB | FL. DSCHIDSCHIGRUT | Glasunov 1892” “ bimaculatus m. det. Luze Type> “Col. et det. A. Fauvel | Coprophilus pentatoma Fvl. | R.I.Sc.N.B. 17.479” “Lectotypus ♂ 1990 C. (Zonoptilus) bimaculatus Luze | det. dr. Tóth L.” “ Coprophilus (Zonyptilus) pentatoma Fauvel, 1897 | det. M. Gildenkov, 2016 ” (IRSNB).

Discussion. The structure of the body and the aedeagus of the lectotype of C. bimaculatus are similar to those of the lectotype of C. pentatoma . The lectotype of C. bimaculatus differs only in having yellow apices of the elytra, which cannot be regarded as a sufficient reason to retain the validity of the species. Thus, the synonymy is established: C. (Z.) pentatoma Fauvel, 1897 = C. (Z.) bimaculatus Luze, 1904 , syn. n.

Remarks. Coprophilus bimaculatus is almost identical in its coloration to C. longicornis and this was apparently the reason why they were considered as synonyms ( Bernhauer, 1908). However, when the lectotype was designated ( Tóth, 1991), C. bimaculatus was accepted as valid species. A detailed discussion of C. bimaculatus as a valid species and illustrations of its aedeagus ( Tóth, 1991), which are allegedly different from the illustrations of the aedeagi of C. pentatoma and C. longicornis , should be regarded as erroneous.

Figures

FIGURES 4 – 5. Coprophilus spp.: 4 — C. (Z.) longicornis Bernhauer, 1903 (holotype, male), body, dorsal view; 5 — C. (Z.) bimaculatus Luze, 1904 (lectotype, male), body, dorsal view. Scale bar: figs. 4, 5 — 1 mm.