Cormocephalus (C.) ungulatus ( Meinert, 1886 )

Schileyko, Arkady A. & Cupul-Magaña, Fabio G., 2021, Cormocephalus (Cormocephalus) guildingii Newport, 1845 (Chilopoda: Scolopendromorpha): a composite description, new samples from Western Mexico and a new species subgroup of Neotropical Cormocephalus (Cormocephalus), Zootaxa 5071 (3), pp. 301-325 : 314-316

publication ID	https://doi.org/ 10.11646/zootaxa.5071.3.1
publication LSID	lsid:zoobank.org:pub:2C1EE869-A61C-4568-9108-5A2397E6D12F
DOI	https://doi.org/10.5281/zenodo.5735532
persistent identifier	https://treatment.plazi.org/id/03AA87D3-FFFA-892B-FF21-DD61FB2AFC03
treatment provided by	Plazi
scientific name	Cormocephalus (C.) ungulatus ( Meinert, 1886 )
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Treatment

Cormocephalus (C.) ungulatus ( Meinert, 1886) View in CoL

Figs 33–38 View FIGURES 30–36 View FIGURES 37–42

Cupipes ungulatus Meinert, 1886: 187 View in CoL ;

Cupipes graecus Meinert, 1887: 123 View in CoL ;

Cupipes ungulatus: Pocock, 1893: 460 View in CoL ;

Cupipes ungulatus: Kraepelin, 1903: 177 View in CoL ;

Cupipes ungulatus: Chamberlin, 1914: 184 View in CoL ;

Cupipes ungulatus: Chamberlin, 1921: 17 View in CoL ;

Cupipes ungulatus: Chamberlin, 1922: 7 View in CoL ;

Cupipes ungulatus: Chamberlin, 1925: 37 View in CoL ;

Cormocephalus (C.) ungulatus: Attems, 1930: 101 View in CoL ;

Cormocephalus (C.) ungulatus: Bücherl, 1941: 300 View in CoL ;

Cormocephalus mundus Chamberlin, 1955: 46 View in CoL ;

Cormocephalus (Cupipes) tingonus Chamberlin, 1957: 31 View in CoL ;

Cormocephalus mundus: Kraus, 1957: 383 View in CoL ;

Cormocephalus tingonus: Bücherl, 1974: 103 View in CoL ;

Cormocephalus ungulatus: Schileyko 2018: 72 View in CoL ;

Cormocephalus ungulatus: Martínez-Muñoz & Perez-Gelabert 2018: 83 View in CoL , 92.

Terra typica: Haiti (= Hispaniola) Island and Pernambuco (Eastern Brazil).

Studied material. Panama, IBISCA project, Colón Province, San Lorenzo Forest , tree canopies, 1 sad. (Rc 7155), 2004, col. I. Tuf. Brazil, Amazonas, Reserva Florestal A. Ducke, near [about 25 km N of] Manaus, 59 59’W, 2 55’S, primary tropical rainforest forest on Terra firme, 1 ad. (Rс 6483), 06-011. III.1998, col. S.I. Golovatsch. GoogleMaps

Description of ad. Rc 6483 [data on sad. Rc 7155 in square brackets where it differs].

Length ca. 30 [15] mm. Coloration in alcohol uniformly yellow [greyish] with head visibly darker [head, body (including sternites) and ultimate legs with small aggregations of dark pigment, Figs 33, 34 View FIGURES 30–36 ]; numerous small setae at all body surface.

Head ( Fig. 35 View FIGURES 30–36 ) with much shortened anteriorly (as long as ½ [1/3] of head), but well-developed [indefinite] paramedian sutures; very short antennae (left 15, right 17 articles [both 17]) hardly extending to the posterior margin of tergite 1 [2] when reflexed.

Forcipular coxosternite ( Figs 36, 33 View FIGURES 30–36 ) with two much shortened posteriorly (as long as 1/2 [1/3] of coxosternite) and converging anteriorly typical longitudinal sutures; transverse suture absent. Tooth-plate with 4 teeth, of these the most lateral and the most median ones are approximately of the same length, being much shorter than 2 remaining teeth; long and pointed trochanteroprefemoral process with one lateral tubercle.

Tergites 1–20 with complete paramedian sutures; tergite 21 ( Fig. 38 View FIGURES 37–42 ), with complete median suture [plus a median depression in the posterior half], only this tergite is marginated laterally.

Sternites 2–20 with complete paramedian sutures disposed in paramedian sulci; sternite 21 ( Fig. 38 View FIGURES 37–42 ) trapeziform, slightly [noticeably] longer than wide, its posterior margin rounded [straight]; sternite 21 with a short, shallow median sulcus / depression in the posterior third.

Legs with 2 short [minute] but clearly recognizable [hardly recognizable] pretarsal accessory spines.

Ultimate LBS ( Fig. 37 View FIGURES 37–42 ): coxal pore field as long as [very slightly longer than] sternite 21, coxopleuron slightly longer than sternite 21, with rounded posterior margin, lacking either any process or spines [the only minute spine at the very inner corner of the coxopleuron].

Ultimate legs ( Figs 37, 38 View FIGURES 37–42 ): prefemur, femur and tibia much flattened dorsally. These articles are each with characteristic dorso-medial longitudinal depression apically; tibia and tarsus 1 somewhat swollen / bulbous ventrally. Prefemur with the only spine at the position of the corner spine ( Fig. 38 View FIGURES 37–42 ) [prefemur ( Fig. 34 View FIGURES 30–36 ) with 3 unusually large dorso-medial spines—2 of them disposed at a remarkably enlarged corner spine—plus 1 ventro-medial and 1 medial one, both of the latter are disposed at the distal margin of the prefemur, forming a transverse row together with the corner spine; no any ventral spines].

Remarks. Adult spm Rc 6483 has been mentioned by Schileyko (2002) as C. brasiliensis Humbert & Saussure, 1870 but the present re-investigation shows that it is C. ungulatus , because it shares with the latter such important peculiarities of the forcipular coxosternite as very short paramedian sutures plus a total absence of a transverse one. This specimen also has no paired dorso-distal “bifid spines” on the ultimate femur (!), which should be diagnostic for C. brasiliensis (see Chamberlin 1914: 183) and being very unusual (if not unique) within Cormocephalus ; Lewis (1989: 1006) wrote that C. brasiliensis “was described on the basis of a defective specimen”.

Kingdom	Animalia
Phylum	Arthropoda
Class	Chilopoda
Order	Scolopendromorpha
Family	Scolopendridae
SubFamily	Scolopendrinae
Genus	Cormocephalus