Cossmannea desvoidyi ( d'Orbigny , 1851)

Gruendel, Joachim, Keupp, Helmut, Lang, Fritz & Nuetzel, Alexander, 2022, Late Jurassic (Upper Kimmeridgian) Heterobranchia (Gastropoda) of the coral-facies of Saal near Kelheim and the viciniy of Nattheim (Germany), Zitteliana 96, pp. 179-221 : 179

Cossmannea desvoidyi ( d'Orbigny, 1851)

Plate 1: figs 1-3

?*1836 - Nerinea grandis Voltz in litt. - Bronn: 549, pl. 6, fig. 1.

*1851 - Nerinea desvoidyi d’Orbigny - d’Orbigny: 107, pl. 261, figs 1-3.

*1858 - Nerinea grandis concava - Quenstedt: 766, pl. 94, fig. 3.

v1881-1884 - Nerinea grandis concava - Quenstedt: 524, pl. 205, fig. 60.

part1901 - Nerinea desvoidyi d’Orb. - Geiger: 283, pl. 11, fig. 5.

1979 - Cossmannea desvoidyi desvoidyi ( d’Orbigny, 1850) - Wieczorek: 321, fig. 11, pl. 7, figs 1, 4; pl. 8, figs 1, 6.

1997 - Cossmannea desvoidyi ( d’Orbigny, 1851) - Fischer and Weber: 42, pl. 11, figs 1-3.

1997 - Cossmannea (Cossmannea) desvoidyi ( d’Orbigny, 1850) - Hägele: 129, fig. p. 129 upper left.

2014 - Cossmannea desvoidyi ( d’Orbigny, 1850) - Kollmann: 354, fig. 2B.

Material.

A specimen from the area near Nattheim (collection Sauerborn) and a specimen from the Quenstedt collection, possibly the specimen figured by Quenstedt (1858, pl. 94, fig. 3) from Nattheim.

Description.

The specimen from the Sauerborn collection is 23.8 cm high. The shell is slender with relatively high whorls. The whorl face is concave and lacks any visible ornament. The suture is distinct but shallow, situated at the lower side of an abapical, rounded bulge forming the transition to the flat, seemingly smooth base. The periphery is situated directly above abapical suture. The aperture is rhomboid with a large, oblique siphonal canal. Two columellar plaits are visible.

Remarks.

According to Fischer and Weber (1997), Cossmannea desvoidyi has a long range from the middle Oxfordian to lower Portlandian. The type specimen is from the middle Oxfordian of France. Considering this long range, Nerinea grandis Voltz in Bronn, 1836 (Portlandian?) and Nerinea grandis concava Quenstedt, 1858 (upper Kimmeridgian) could represent C. desvoidyi . If true, then Nerinea grandis would be the oldest available name for this species.

The identity of C. desvoidyi is unclear, especially regarding the plaits, because there are conflicting reports in the literature: According to Geiger (1901), Cox (1947), and Fischer and Weber (1997), C. desvoidyi has a single columellar and a single palatal plait. In the present specimen from Nattheim, two columellar plaits are visible. The outer lip is not sufficiently well-preserved to show or exclude the presence of palatal plaits. Bronn (1836) and Kollmann (2014) mention the presence of only a single columellar plait. It is unclear whether these differences are results of variability, preservation, ontogenetic change, or whether two or more species are present.

There are also contradictory statements about the morphology of the aperture of C. desvoidyi . Kollmann (2014) reported that a siphonal canal is absent in specimens from the type locality of St. Mihiel. However, Fischer and Weber (1997: pl. 11, fig. 1) figured a specimen from the same locality that has a siphonal canal. The present specimen from Nattheim (Sauerborn collection) has a very distinct oblique canal. Furthermore, this species is generally said to lack ornamentation, but the illustration by Geiger (1901: pl.11, fig. 5) shows numerous spiral cords covering the entire whorls, which is, however, not mentioned in Geiger‘s (1901) description of this species.

The present contribution cannot solve these problems. We assume that the upper Kimmeridgian specimens described herein represent the same species as the middle Oxfordian holotype of C. desvoidyi from France. In order to avoid nomenclatorial confusion, we include only the reference to the type material and to Kimmeridgian material in the synonymy and chresonymy list.

Figures

Plate 1. (1 - 3) Cossmannea desvoidyi (d'Orbigny, 1851). (1 - 2) collection Sauerborn, Nattheim area; (1) lateral view, height 23.8 cm; (2) last whorls with aperture, height 8 cm. (3) University Tuebingen, collection Quenstedt), Nattheim, lateral view, height 130 mm (original of Quenstedt 1881 - 1884, pl. 205, 60). (4 - 6) Aptyxiella planata (Quenstedt, 1858), SNSB-BSPG 2021 XV 1, Nattheim-Rinderberg; (4 - 5) lateral views, height 70 mm; (6) aperture, height of detail 70 mm. (7) Aptyxiella quenstedti Geiger, 1901, Nattheim, lateral view (copy from Geiger 1901, pl. 11, fig. 1).