Dalmatoreicheia maderi, Bulirsch, Petr & Guéorguiev, Borislav V., 2008

Bulirsch, Petr & Guéorguiev, Borislav V., 2008, Taxonomic studies of the Balkan Reicheina (Coleoptera: Carabidae: Clivinini), with a review of the Albanian species, Zootaxa 1679, pp. 39-53 : 40-42

Dalmatoreicheia maderi sp. nov.

( Figs 1 , 4 , 7 , 9 , 12 , 16 )

Type material. Holotype ɗ labelled: “Kruja, Al­ban., Mader”; “ Holotype, Dalmatoreicheia maderi sp. n., Bulirsch et Guéorguiev des. 2006” ( NMW).

Description. Body rusty brown, elytra slightly darker, antennae and mouthparts rusty yellow, legs light rusty red, front legs slightly darker ( Fig. 1 ). Length 3.20 mm.

Head. Narrow, long, anterior margin of clypeus moderately and regularly emarginated; frons with facial furrows shortened, shallow and broad. Impressions on clypeus oblique, broad, shallow, interrupted in middle. Eyes absent, genae with median part long, slightly vaulted, their hind angles distinct. Vertex rather shiny, with moderately reticulated microsculpture. Left antenna length 1.20 mm (right antenna missing except antennomere 1); antennomere 2 just longer than 3 and 4 combined, antennomeres 6–10 slightly longer than wide. Mandibles rather long, their apical part moderately curved. Terminal maxillary palpomeres moderately long, with apex slightly narrowed, not acicular.

Pronotum. Moderately convex, microsculpture distinct, reticulation regularly diffused. Sides regularly and moderately rounded, attenuated posteriorly; maximum width before middle; posterior angles very broadly rounded. Reflexed lateral margin entire, extended from short, sharp, rather protruding anterior angles to base as very distinct prebasal groove. Median line deep, distinctly impressed almost towards prebasal groove; anterior transverse impression shallow, slightly deeper laterally. Episterna just visible from above in extreme apical part. Lateral channel on each side with 2 setiferous punctures, anterior pair slightly separated from lateral margin; neither sublateral nor submedial discal setae. Ratio length: width 1.08; pronotum 1.60 times as broad as head.

Legs. Protibia with apical spine bent outwards in dorsal view, apical spur of almost equal length to spine, slightly curved. Lower marginal tooth large, prominent, upper one distinct, sharp. Hind tarsi length 0.50 mm; first tarsomere elongated.

Elytra. Convex, disc flattened, outline almost oval, maximum width at middle. Base distinctly bordered, strongly sloping, without distinct granules; prescutellar setiferous punctures isolated; suture broadly depressed; humeri very broadly rounded, not protruding. Lateral channel very wide, reflexed lateral margin with about 30 denticles; 6–8 humeral denticles sharp, lateral and especially apical ones much finer, just recognizable almost up to apex with slightly protruded suture. Striae 1–5 distinct, slightly punctate, striae 6–7 finer, moderately punctate; all striae strongly diminish apically: first striae in apical fifth, lateral ones in apical third; apically with only traces of fine punctures. Striae 1–3 distinct almost up to base, striae 4–7 slightly to moderately shortened. Intervals 1–5 in basal part slightly convex, lateral ones flattened. Intervals 2–7 with rows of setae; punctures in first intervals broader than striae punctures. Ratio length: width 1.71; ratio elytra: pronotum length 2.11; ratio elytra: pronotum width 1.35.

Venter. Last visible ventral segment in male with slight reticulate microsculpture in apical half.

Male genital armature. Ninth postabdominal sternum (urite) as in Fig. 4 ; ovate, short, widely rounded in basal part and briefly rounded in apical part; sclerites fused. Median lobe as in Figs 7 , 9 ; arcuate in lateral view, apical part wider than basal part; apical orifices straight, basal orifice concave; internal sac situated in subapical part; copulatory piece forming inside four more or less visible lamellae; median lobe in ventral view with both apex and basis slightly bent to left side. Parameres dissimilar in shape and size as in Fig. 12 ; left one smaller, strongly bent at about middle, apex with one short, distinctly removed preapical seta, and two long apical setae; right paramere massive, with two long apical setae.

Female genital armature. Unknown.

Differential diagnosis. D. maderi sp. nov. can be distinguished from D. janaki Magrini & Bulirsch, 2005 , the single hitherto known species of this genus, by its narrow pronotum (ratio length: width 1.08 versus 0.96); by the strongly oblique base of the elytra and indistinct humeri and by the presence of rows of setae in intervals 2–7 (versus in intervals 2, 3, 5, 7 only).

Etymology. The species is named in honor of the collector, the Austrian coleopterologist and biospeleologist Leopold Mader (1886–1961).

Type locality. Krujë is a well­known historical town, 32 km north of the capital Tiranë, in the central­east part of Albania. The town is situated on an isolated spur of a limestone mountain­wall of the massif of Sari­ Salltiku, part of the Krujë Range (altitude ca. 550 m above sea level). The exact place of finding is unknown.

Remarks. We suppose that the holotype of D. maderi sp. nov. had been collected before 1922, judging by the year of printing of Mader´s paper in the Wiener Entomologische Zeitung ( Mader 1921). There the author described Epierus krujanensis Mader, 1921 (currently junior synonym to the histerid beetle Carcinops pumilio (Erichson, 1834)) from a type locality identical with our new species.

The genus Dalmatoreicheia is known only from two specimens: a female of D. janaki from Croatia, the type species, and a male of D. maderi sp. nov. It is necessary to find further material to verify the congeneric status of both taxa.

Figures

FIGURE 16. Map of Albania with localities of species. Star: Dalmatoreicheia maderi sp. nov.; black circle: Reicheadella bischoffi; reverse triangle: R. smetanai sp. nov.

FIGURES 1 – 3. Habitus of type species (length in parentheses behind the name). Fig. 1, Dalmatoreicheia maderi sp. nov., (3.20 mm), holotype; Fig. 2, Reicheadella bischoffi, (2.80 mm), lectotype; Fig. 3, Reicheadella smetanai sp. nov., (2.05 mm), holotype.

FIGURES 4 – 6. Ventral view of postabdominal sternum IX (urite). Fig. 4, Dalmatoreicheia maderi sp. nov., holotype; Fig. 5. Reicheadella bischoffi, lectotype; Fig. 6, Reicheadella corcyrea, male labelled “ Gasturi, Korfu 2. IV. (19) 29. Beier ” (NMW). Scale bar = 0.1 mm.

FIGURES 7 – 8. Left lateral view of median lobe of aedeagus. Fig. 7, Dalmatoreicheia maderi sp. nov., holotype; Fig. 8. Reicheadella bischoffi, lectotype. Scale bar = 0.1 mm.

FIGURES 9 – 11. Ventral view of median lobe of aedeagus. Fig. 9, Dalmatoreicheia maderi sp. nov., holotype; Fig. 10, Reicheadella bischoffi, lectotype; Fig. 11, Reicheadella corcyrea, male labelled “ Gasturi, Korfu 2. IV. (19) 29. Beier ” (NMW). Scale bar = 0.1 mm.

FIGURES 12 – 14. Ventral view of parameres. Fig. 12, Dalmatoreicheia maderi sp. nov., holotype; Fig. 13, Reicheadella bischoffi, lectotype; Fig. 14, Reicheadella corcyrea, male labelled “ Gasturi, Korfu 2. IV. (19) 29. Beier ” (NMW). Scale bar = 0.1 mm.