Diplommatina javana Möllendorff, 1897

Hausdorf, Bernhard, 2017, Revision of the Diplommatinidae (Gastropoda: Cyclophoroidea) from Java, Zootaxa 4312 (2), pp. 201-245 : 211

Diplommatina javana Möllendorff, 1897

( Figs 4 , 10 , 24 ; Table 1)

Diplommatina (Sinica) javana Möllendorff, 1897: 97 (JaVa) . Kobelt 1902: 465. Zilch 1953: 31, pl. 12 fig. 178.

Diplommatina (Sinica) javana orientalis RenSch, 1931: 387 ( FloreS: Geli Moetoe Volkan [1300 m]).

Diplommatina javana — Van Benthem Jutting 1948: 599, fig. 55. Vermeulen & Whitten 1998: 50, 138, fig. 28.

Diplommatina sulcicollis —Heryanto 2008: 362 (partly, not Möllendorff 1897). Heryanto 2009: 332 (partly, not Möllendorff 1897). Heryanto 2012: 196 (not Möllendorff 1897).

Palaina nubigena —Heryanto 2008: 362 (partly, not Möllendorff 1897). Heryanto 2009: 332 (partly, not Möllendorff 1897).

Diagnosis. Diplommatina javana is characterized by a closely ribbed, small (shell height 2.65–3.2 mm), sinistral shell with a convex last whorl without a spiral depression, a slightly expanded peristome and with a spiral palatal fold close to the suture ventrolaterally right inside the last whorl.

Shell ( Figs 4 , 24 ; Table 1). Sinistral; ovoid with conical early whorls and a narrowed last whorl; with 5 whorls separated by a deep suture; protoconch smooth; teleoconch with oblique ribs (58–80 ribs on the penultimate whorl; approximately the same number on the last whorl); locally with spiral striae between the ribs; corneous; subtranslucent; last whorl rounded, with an indistinct ventral constriction; aperture subangular, angular at the transition from columellar to basal side; upper insertion of the peristome distinctly ascending towards aperture; insertions of the peristome connected by a weak callus; peristome expanded, thickened, doubled only towards the columella; aperture with a distinct columellar lamella; with a short spiral palatal fold close to the suture ventrolaterally right inside the last whorl and with a radial palatal fold ventrally close to the suture; umbilicus closed; operculum thin, corneous, ovoid, with an arcuate ridge on the inner surface, paucispiral. Shell height 2.65– 3.2 mm, shell diameter 1.55–2.0 mm.

Remarks. Rensch (1931) compared samples of D. javana from Bali and Flores with a sample from Mt. Cikuray, which he considered to represent the nominotypical subspecies of D. javana . Because the specimens from Mt. Cikuray had 28–34 ribs on the penultimate whorl, whereas the specimens from Bali and Flores had 65–75 ribs, Rensch (1931) separated the latter as D. javana orientalis Rensch, 1931 . Actually, the sample from Mt. Cikuray did not represent D. javana , but Diplommatina kakenca n. sp., whereas the samples from Bali and Flores do not differ from the true D. javana (see also Vermeulen & Whitten 1998).

Habitat. In leaf litter and on deadwood in primary and secondary forests on volcanic soils at 500–2700 m a.s.l.

Type material. Indonesia, Java (lectotype of D. javana [designated by Zilch 1953] SMF 105202; paralectotypes of D. javana SMF 105203/4).

Other material. Indonesia, Java (SMF 192585/4).—West Java: West Bandung, Mt. Tilu, 1900 m a.s.l., 7°06'S 107°23'E (SMF 105205/2, vouchers to Zilch 1953); Talun near Bandung, 800 m a.s.l., 7°07'S 107°34'E (ZMA 68363/1, vouchers to van Benthem Jutting 1948); Garut, summit of Mt. Papandayan, 2600 m a.s.l., 7°19'48''S 107°43'16''E (ZMA 68364/2, vouchers to van Benthem Jutting 1948); Garut, Mt. Papandayan, 2300 m a.s.l., 7°16'S 107°44'E (SMF 105206/1, vouchers to Zilch 1953); Garut, Mt. Cikuray, 1900 m a.s.l., 7°18'S 107°52'E (SMF 105204/4, vouchers to Zilch 1953); Garut, Mt. Cikuray, 2700 m a.s.l., 7°19'S 107°52'E (van Benthem Jutting 1948); Majalengka, Argamukti, Mt. Ciremai National Park, 6°54'S 108°22'E (MZB 13255/1, voucher to Heryanto 2008, 2009 [as P. nubigena ]; MZB 13261/52, vouchers to Heryanto 2008, 2009 [as D. sulcicollis ]); Kuningan, Palutungan, 6°55'S 108°24'E (MZB 17971/1); Kuningan, Mt. Ciremai, Kiara Bolong, 2320 m a.s.l., 6°55'S 108°24'E (BDJ/2).—Central Java: Purbalingga, Karangreja village, Mt. Slamet, 1405 m a.s.l., 7°13'S 109°15'E (MZB 15509/1, 15519/1, vouchers to Heryanto 2012 [as D. sulcicollis ]).

Distribution ( Fig. 10 ). Diplommatina javana is known from Java, Bali and Flores ( Rensch 1931; Vermeulen & Whitten 1998).

Figures

FIGURES 4 – 8. Opercula of Diplommatinidae. 4. Diplommatina javana Möllendorff, 1897, outer and inner Surface of opercula, WeSt JaVa: Majalengka, Argamukti, Mt. Ciremai National Park (MZB 13261). 5. Diplommatina auriculata Möllendorff, 1897, inner Surface of operculum, WeSt JaVa: Majalengka, Argamukti, Mt. Ciremai National Park (MZB 13258). 6. Palaina gedeana Möllendorff, 1897, outer and inner Surface of opercula, WeSt JaVa: Majalengka, Argamukti, Mt. Ciremai National Park (MZB 13244). 7. Palaina nubigena Möllendorff, 1897, outer Surface of operculum, WeSt JaVa: Cianjur, Kandang Badak (ZMA 68468). 8. Arinia yanseni n. sp., inner Surface of operculum, EaSt JaVa: Rembang, Sumber PacuSen (MZB 18882). Scale bar = 0.07 mm for FigS 3 – 6, 0.1 mm for Fig. 7.

FIGURES 10 – 14. DiStribution of Diplommatinidae in JaVa. 10. Diplommatina planicollis Möllendorff, 1897, Diplommatina heteroglypha Van Benthem Jutting, 1948 and Diplommatina javana Möllendorff, 1897. 11. Diplommatina halimunensis n. sp., Diplommatina kakenca n. sp. and Diplommatina heryantoi n. sp. 12. Diplommatina perpusilla Möllendorff, 1897 and Diplommatina ristiae n. sp. 13. Diplommatina sulcicollis Möllendorff, 1897 and Diplommatina cyclostoma Möllendorff, 1897. 14. Diplommatina calcarata Möllendorff, 1897 and Diplommatina tetragonostoma Möllendorff, 1897.

FIGURES 23 – 25. Diplommatina Spp. from JaVa. 23. Diplommatina heteroglypha Van Benthem Jutting, 1948, EaSt JaVa: Banyuwangi, Mt. Ijen, Merapi Ungup-ungup (paratype ZMA 135947). 24. Diplommatina javana Möllendorff, 1897, JaVa (lectotype SMF 105202). 25. Diplommatina cyclostoma Möllendorff, 1897, WeSt JaVa: Cikuray (lectotype SMF 105214). Scale bar = 1.0 mm.