Drastria picta picta ( Christoph, 1877 )

Yu, Alexey & Korb, Stanislav K., 2019, A revision of the genus Drasteria of Central Asia and Kazakhstan with special attention to the adjacent areas (Lepidoptera: Erebidae), Zootaxa 4673 (1), pp. 1-104 : 55-57

Drastria picta picta ( Christoph, 1877)

(= D. radapicta ( Staudinger, 1901) , = D. austera ( John, 1921) , syn. n.) ( Map 19 ) ( Figs. 142–153 (imago), 248, 249 (male genitalia), 286, 293 (female genitalia))

Christoph, 1877: 257; Tab. 7, fig. 28 ( Leucanitis cailino var. picta ). Type locality: “bei Krasnowodsk” (by the original description).

Type material. Lectotype male (designated here in accordance with ICZN Art. 74) ( Figs. 142–144 ), labelled: “Origin.” (pink paper, printed), “ex coll. 2/3 | Staudinger” (white paper, printed with handwritten numbers), “Krasnow. | Christ.” (yellow paper, handwritten), “ LECTOTYPUS ♀ | Leucanitis cailino var. picta | Christoph, 1877 | Horae Soc. ent. Ross. 12 (3): 257, (4) pl. 7, f. 28 | S. Korb et A. Matov det. 2019” (red paper, printed) ( ZMHU) . Paralectotypes: 2 females ( ZMHU) .

= radapicta Staudinger in Staudinger et Rebel, 1901: 243 ( Leucanitis Picta (Cailino var.) v. Radapicta). Type locality: “Ross. mer. or. (Narün et Kirgh.)” (by the original description); “Narün” (by the lectotype designation, designated here).

Type material. Lectotype male (designated here in accordance with ICZN Art. 74) ( Figs. 145–147 ), labelled: “Origin” (pink paper, printed), “ex coll. 1/8 | Staudinger” (white paper, printed with handwritten numbers), “Narün | Henke” (green paper, handwritten), “ LECTOTYPUS | Leucanitis picta radapicta | Staudinger, Rebel, 1901 | Cat. Lep. Faunengeb. 1: 243 | Korb, Matov, Volynkin des. | 2016, Feb. 1-4 ” (red paper, printed) ( ZMHU) . Paralectotypes: 5 males, 2 females ( ZMHU) .

= austera John, 1921: 44 ( Leucanitis austera ). Type locality: “ Persia, Beludchistan: Megas, Kooch-y-Birk” (by original description).

Type material: Holotype by monotypy, female ( Figs. 151–153 ), labelled: “sp. typ. | austera | John” (white paper, handwritten); “Мегас, гКухиБирик, | БелудЖ; ПерсиЯ | Зарудный. 12 II 01 ” < Megas , Kukhi-Birik Mt. , Beludzhistan, Persia, Zarudny 12.ii.1901> (white paper, printed) ( ZISP).

Remark. There are no differences in the genitalia and wing pattern of the type specimens of the taxa picta , radapicta and austera , thus, we treated them here as synonyms. All of them have very characteristic external feature: the discal spot on the hindwing underside (see the “Differential diagnosis” of the previous species). The female genitalia have the same size of the sclerotized plate in the ostium bursae, which is two times shorter in D. pseudopicta Matov et Korb , sp. n. In the male genitalia this species differs from D. pseudopicta by the absence of the thin diverticulum between two main lobes of vesica and by the shape of these lobes.

Alphéraky, 1882: 93 [Khorgosse]; Butler, 1892: 316 [Krasnow.[odsk]]; Staudinger, Rebel, 1901: 243 [Ili]; Kusnezov, 1908: 112 [Agespe, Kulandy]; John, 1910: 605 [Emba, Tschelkar, Malyje Barsuki, Perovsk, Dshulek]; Hampson, 1913 b: 282 [Ili]; Warren W., 1913: 342 [Ili]; Culot, 1913: 184 [la Russie méridionale orientale]; Kuznetzov, 1960: 87 [Ay-Dere]; Stshetkn, 1956: 135 [Kara-Dum and Kashka-Dum sands]; Daricheva, 1962: 84 [Krasnovodsk, Askhabad]; Daricheva, 1965: 57 [Krasnovodsk; Ai-Dere; Askhabad; Bobozo near Askhabad]; Stshetkin, 1965: 167 [Kara-Dum and Kashka-Kum (Mt. Buritau) sands, Khalka-Kul lake]; Sinadsky, 1968: 80 [ Kazakhstan]; Sviridov, 1971: 47 [Badkhyz: Er-Oylan-Duz lake]; Shek, 1972: 143 [Alma-Ata Prov.]; Aibasov, 1974: 123 [Ustyurt plateau: Karynzharykskaya depression]; Shekh, 1975: 412 [Alma-Ata Prov]; Ronkay, 1983: 236 [Bajanchongor aimak: Oase Echin gol, cca 90 km NO Von Grenzposten Caganbulag, 950 m; Gobi Altaj aimak: Zachuj Gobi, 10 km N von Chatan chajchan Gebirge, 1l 50 m; Südgobiaimak: Ostrand von Zöölön ul Gebirge, 58 km WSW von Somon Bajan-dalaj, 1500 m; Uburchangaj aimak: am halben Weg zwischen Somon Bajanleg und Somon Bulgan, cca 130 km OSO von Bajanleg, 1150 m]; Nekrasov et al., 1988: 51 [Osh]; Poole, 1989: 329; Beck, 1996: 25; Milko, 1996: 258 [Regions close to Fergana (including southern slopes of Chatkalsky and Fergansky Mts. and northern slopes of Alai and Turkestansky Mts.)]; Klyuchko, Kreuzberg, 1998: 26 [Nuratinsky Nature Reserve]; Lehmann et al., 1998: 526 [Ili area, Tschu river, Mujunkum, 200m; Ili river, Bakanas, 400m; Ili area, Charyn river, 400m]; Poltavsky et al., 1998: 108 [Tshilmamed-Kum, Imam-Baba, Bairam-Ali, Kizyl-Arvat, Tschardzhou, Badkhyz, Ai-Dere, Yaskhan, Repetek, Dardja, Ashkhabad]; Ivinskis, Miatleuski, 1999: 204 [Repetek, Ashkhabad, Kov-Ata]; Anikin et al., 2000: 332 [Volgograd, Saratov]; Poltavsky, Ilyina, 2002: 17 [Kumtor-Kale, Derbent]; Poltavsky, Nekrasov, 2002: 26 [Daghestan Republic]; Goater et al., 2003: 78 [South Russia: Saratov and Volgograd Districts and Kazakhstan: Botkul, Urda, Kandagash, Bisen and Dzhaylau. Outside Europe, in Transcaspia, Turkestan to Central Asia]; Lehmann, Bergmann, 2005: 20 [Issyk Kul depression, Kirgizsky Mts., Alai Mts.]; Kononenko, 2010: 89 [ Kazakhstan]; Gorbunov et al., 2011: 76 [Ustyurt Nature Reserve, Bostankum sands, 40 km NE Zhanaozen, 43°39’ N, 53°08’ E; Ustyurt Nature reserve, Sengirkum sands, 43°47’ N, 53°34’ E; Ustyurt Nature reserve, Besokty dwell, 18 km E Senek, 43°20’ N, 53°36’ E; Northern slope of Ustyurt Nature reserve, 43°24’ N, 54°33’ E; Ustyurt Nature Reserve, S of Karashek Mt. 18 km N of Akkuduk, 43°06’54” N, 54°11’40” E; Ustyurt Nature reserve, Kendyrli dwell, 42°57’ N, 54°41’ E; Ustyurt Nature reserve, Kokesem dwell, 43° 10’ N, 54°53’ E; Ustyurt Nature reserve, Kendyrli-sor 15 km W of Kendyrli, 42°59’ N, 54°32’ E; Ustyurt Nature reserve, Zhomarta, 13 km SE Akkuduk, 42°55’ N, 54°15’ E; Ustyurt Nature reserve, Saksorka dwell, 30 km S of Akkuduk, 42°42’ N, 54°06’ E; Ustyurt Nature reserve, Onere dwell, 42°36’ N, 54°08’ E]; Korb, 2013: 28 [3 km N of Kapchagai, Ili River Valley, N 43 55.861 E 77 05.513, 574 m; 35 km N of Bakanas, Ili River Valley, N 44 53.940 E 75 53.479, 377 m; Singing Dune in Altyn Emel Nature Reserve, eastern border of Kapchagai Reservoir, Ili River Valley, N 43 50.273 E 78 35.759, 467 m; Bridge through Ili River 22 km SE of Koktal, Ili River Valley, N 43 58.004 E 79 35.905, 600 m]; Matov, Anikin, 2015: 45 [Astrakhan Prov., Stepnoe; Saratov Prov., Khvalyn distr., 5 km W Khvalynsk]; Shovkoon, Trofimova, 2016: 607 [Aktobe Prov., near Kompressorniy vill. N 48°30’; E 59°19’; Aktobe Prov., N 47°59’ E 59°24’; Aktobe Prov., Bolshie Barsuki sands, N 46°58’; E 59°13’; Aktobe Prov., Shagyrai Plateau, N 46°52’; E 58°02’; Aktobe Prov., Shagyrai Plateau, N 46°50’; E 57°47’; Aktobe Prov., 35 km S Begembet vill., N 46°48’; E 59°09’; Aktobe Prov., Shagyrai Plateau, N 46°44’; E 57°30’; Kyzyl-Orda Prov., Karatup penin, Aral lake, N 46°20’; E 59°43’; Aktobe Prov., near Bozoe vill., sands, N 46°17’; E 58°56’; Mangistau Prov., Mangistau Mts., N 44°18’; E 51°35’; Mangistau Prov., Ustyurt Plateau, N 43°57’; E 53°45’].

Habitat ( Fig. 303 ). Deserts and semideserts, prefer sands. Elevation: 200– 500 m.

Distribution. Southern part of European Russia (Astrakhan, Saratov, Orenburg, Volgograd Prov.), deserts of Central Asia.

Material examined (in total: 155 specimens). Russia. 6 ♂, 2 ♀, Narün (lectotype and paralectotype of radapicta ) ( ZMHU) ; 1 ♂, 2.05.2010, Astrakhan Prov., Dosang , desert, dunes, leg. S.K. Korb ( ZISP) . China. 1 ♀, 04.1879, Khorgos , leg. Alph [eraky], ex coll. Grand Duke Nikolai Mikhailovich ( ZISP) ; 1 ♀, Kuldja merid., leg. Alph [eraky], ex coll. Grand Duke Nikolai Mikhailovich ( ZISP) ; 1 ♀, Lob-noor , leg. Przewalsky, ex coll. Grand Duke Nikolai Mikhailovich ( ZISP) . Iran. 1 ♀, 12.02.1901, Kukhi-Birik Mt., Beludzhistan, Persia , (holotype of austera ) ( ZISP) . Kazakhstan. 1 ♀, 2.07.[?], Charyn , leg. unknown, ex coll. O. John ( ZISP) ; 1 ♂, 20.05.[?], Dzhulek , leg. unknown, ex coll. Mus. Acad. Petrop. ( ZISP) ; 1 ♂, 29.05.1899, Turgaisk Prov. , leg. Sushkin ( ZISP) ; 1 ♂, 3 ♀, 25.05.1907, Turgaisk Prov., Bolshie Barsuki desert, Chelkar lake , leg. Volman ( ZISP) ; 1 ♂, 1 ♀, 1908, Syr-Darja, Perovsk , leg. Nikolsky, ex coll. O. John ( ZISP) ; 8 ♂, 3 ♀, 21.04– 13.05.1908, Syr-Darja, St. Baigakum , leg. Malyschew, ex coll. O. John ( ZISP) ; 1 ♂, 18.06.1908, Turgaisk Prov., Chelkar env, leg. N. Androsov ( ZISP) ; 22 ♂, 8 ♀, 8– 29.07.1908, Turgaisk Prov., Malye Barsuki near Kara-Chokat , leg. N. Androsov, ex coll. O. John ( ZISP) ; 3 ♂, 1 ♀, 26.04.1909, 17.07.1910, Syr-Darja, Dshulek , leg. Koshantschikoff, ex coll. O. John ( ZISP) ; 7 ♂, 2 ♀, 1– 9.05.1909, Syr-Darja, Aj-Darle , leg. Koshantschikoff, ex coll. O. John ( ZISP) ; 19 ♂, 5 ♀, 15.05.1910, prov. Syr-Darja, deser. Mujun-Kum, lac. Kargaly-Kul , leg. A. Golbeck, slide N Matov 0552 ( ZISP) ; 2 ♂, 1 ♀, 19.05.1910, Muyun-Kum sands, 5 km S of Karagaly-Kul, leg. A. Golbeck, ex coll. O. John ( ZISP) ; 5 ♂, 4.06.- 9.07.1911, Syr- Daria, Baigacum , leg. Koshantschikoff, ex coll. O. John ( ZISP) ; 1 ♂, 15.08.1930, Priaral Karakum, Baigana-sor , leg. E. Luppova ( ZISP) ; 1 ♂, 21.06.2010, Karaganda Province, Moiyunkum Sands at Sary-Su River , 46º12’ N 67º06’ E, leg. P. Gorbunov ( ZISP) ; 3 ♂, 2– 3.05.2018, Akkergeshen , 47°18’9”N 54°23’22”E, leg. A. Samus ( SKK) ; 3 ♂, 2 ♀, 10– 16.06.2018, Aidarly , 43.982675°N, 79.587779°E, leg. Litovtsev ( SKK) . Kyrghyzstan . 1 ♀, Naryn , leg. unknown, ex coll. O. John ( ZISP) . Mongolia. 2 ♂, 1 ♀, 12.07– 15.08.1981, Bayan-Khongorski aimak, 140 km S of Shine-Dzhinsta, Ekhin-Gol , leg. А. Lvovsky ( ZISP) ; 1 ♂, 1 ♀, 7.07.1981, South Gobi aimak, Tost-Ula moun- tains, leg. А. Lvovsky ( ZISP) . Russia. 6 ♂, 2 ♀, Narün ( ZMHU) ; 8 ♂, 1 ♀, 2.05.2010, Astrakhan Prov., Dosang , desert, dunes, leg. S.K. Korb ( SKK, CKO) . Turkmenistan. 3 ♀, Krasnow. [odsk] (lectotype and paralectotypes) ( ZMHU) ; 3 ♀, 15.04–23.05.[18]91, Aidara , leg. Eylandt, ex coll. Grand Duke Nikolai Mikhailovich, slide N 6468 (Ryabov) ( ZISP) ; 1 ♀, [19]03, Transcaspian Prov. , leg. Svetlovsly ( ZISP) ; 1 ♀, 4–5.1937, Repetek , leg. Lavrov ( ZISP) ; 1 ♂, 1951, Uzboi , SW Burgun, leg. Schteinberg ( ZISP) ; 1 ♀, 4.06.1954, Askhabad vic., leg. Potopolsky, slide N Matov 0546 ( ZISP) ; 1 ♀, 13– 22.05.1955, Aschabad circ., leg. W. Potopolskij ( ZISP) . Uzbekistan. 1 ♂, Russky Turkestan , [leg. Fedchenko], ex coll Erschov ( ZISP) ; 1 ♀, 24.05. 1972, 140 km NW Shafrikan, Zhamansai , leg. М. Falkovich ( ZISP) .

Figures

MAP 19. Distribution map of Drastria picta (Christoph, 1877).

FIGURES 127–147. Drasteria specimens. 127–129. D saisani (Staudinger, 1882), Khosrov Nature Reserve, Armenia (127), Kekemeren river valley, Kyrgyzstan (128), Chulak Gorge, Dzhungarian Alatau Mts., Kazakhstan (129). 130–134. D. chinensis (Alphéraky, 1892), lectotype (130: upperside; 131: underside; 132: labels), Mongolia (133), Panfilov env., Kazakhstan (134). 135. D. catocalis perunovi Volynkin et Matov, 2012, holotype. 136–138. D. catocalis catocalis (Staudinger, 1882), lectotype (136: upperside; 137: underside; 138: labels). 139–141. D. pseudopicta Matov et Korb, sp.n., holotype (139, 140) (Dosang, Astrakhan Prov., Russia) and paratype (141) (Kapchagai, Kazakhstan). 142–147. D. picta (Christoph, 1877), lectotype (142: upperside; 143: underside; 144: labels); lectotype of radapicta (145: upperside; 146: underside; 147: labels).

FIGURES 148–165. Drasteria specimens. 148–153. D. picta (Christoph, 1877), Krasnowodsk, Turkmenistan (148), Sauskan sands, Kazakhstan (149), Aidarly, Kazakhstan (150), holotype of austera John, 1917 (151: upperside; 152: underside; 153: labels). 154–159. D. sesquistria (Eversmann, 1854), lectotype (154: upperside; 155: underside; 156: labels), Ili river valley, Kazakhstan (157), Aidarly, Kazakhstan (158), Sauskan sands, Kazakhstan (159). 160–165. D. aberrans (Staudinger, 1888), lectotype (160: upperside; 161: underside; 162: labels), Kapchagai, Kazakhstan (163), Baian Toroi, Mongolia (164), Aidarly, Kazakhstan (165).

FIGURES 297–304. Drasteria habitats. 297. 25 km N of Panfilov, Dzhungarian Alatau, Kazakhstan (one of the authors in the collecting trip 2010). 298. Ili river valley near Koktal bridge, Almaty Prov., Kazakhstan (Russian Entomological Society and Institute of Zoology of Kazakhstan collecting trip, 2010). 299. Sands near Dosang environs, Astrakhan Prov., Russia. 300. Saksaul forest near Bakanas, Almaty Prov., Kazakhstan. 301. Chulak gorge, foothills of Dzhungarian Alatau Mts. in the Altyn-Emel Nature Reserve, Kazakhstan. 302. Ili river shore near Singing Dunes, Altyn-Emel Nature Reserve, Kazakhstan. 303. Katutau mountains, Altyn-Emel Nature Reserve, Kazakhstan. 304. Koram environs, northern slopes of Transilian Alatau Mts., Kazakhstan. Photos: S.K. Korb.