Ectopioglossa sudanensis Selis, 2023

Selis, Marco, 2023, A new Afrotropical species of Ectopioglossa Perkins (Hymenoptera: Vespidae: Eumeninae), Zootaxa 5318 (2), pp. 268-274 : 271-273

Ectopioglossa sudanensis Selis , sp. nov.

( Fig. 2 )

Diagnosis. Recognized from other Afrotropical Ectopioglossa by the combination of following characters: clypeus strongly convex in lateral view, medially depressed in apical half; F11 small and digitiform, not reaching base of F9 ( Fig. 2E ); pronotum with blunt but distinct oblique humeral carina, separating dorsal face from depressed lateral faces; lateral carina of propodeum sinuous above propodeal valvula, not forming a sharp triangular tooth ( Fig. 2F ); T1 about 3× as long as apically wide ( Fig. 2G ); S1 with narrow basal part present, covered with weak and irregular transverse striae; pronotum with large sparse punctures, leaving interspaces at least as wide as puncture diameter ( Fig. 2D ); dorsal faces of propodeum microreticulate; head and mesosoma with long silvery setae, metasoma with shorter and denser pubescence; entirely dark coloration with very reduced ivory markings on head, pronotum, propodeum, T1–T2 and legs; wings infuscate along costal margin.

Material examined. HOLOTYPE: ♁, labeled “Kagelu, Equatoria / Sudan 8-II-39 / J G Myers 10 394 // Ectopioglossa sudanensis / HOLOTYPUS ♁ / Det. Marco Selis [red label]” ( MSNVE).

Description. Male holotype. Body length 8.5 mm; fore wing length 6 mm.

Head 1.15× as wide as long in frontal view ( Fig. 2C ). Clypeus in frontal view as long as wide, semicircularly emarginate at apex, apical emargination about 0.5× as wide as interantennal distance and 2× as wide as deep, apical angles triangular and apically rounded and bearing short blunt carinae curving out, the carinae continued by rounded folds of the clypeus delimiting a median depressed area; clypeus in lateral view strongly convex in basal third, then flattened to the apex. Interantennal space almost evenly convex, with a weak indication of median longitudinal ridge. Distance from posterior ocellus to occipital margin 1.9× as long as the distance from posterior ocellus to inner eye margin; ocelli disposed as an obtuse triangle, distance between posterior ocelli 2.5× as long as distance between anterior ocellus and posterior ocellus. Gena 0.5× as wide as eye at bottom of ocular sinus; occipital carina complete, shortly lamellate on sides, almost evenly curved from vertex to mandible. Scape 3.5× as long as apically wide; F1 2× as long as wide and 1.4× as long as F2; F2–F9 longer than wide, becoming proportionally shorter, convex on dorsal side; F10 small and subquadrate, barely visible; F11 small and digitiform with rounded apex, very weakly curved in lateral view, not reaching base of F9; tyloids barely indicated on F8–F9, linear and parallel sided ( Fig. 2E ). Mandible short and robust, with five triangular pointed teeth. Mesosoma in dorsal view elongate, 1.6× as long as wide; forming a single even curve from pronotum to apex of propodeum in lateral view. Pronotum in dorsal view with weakly converging sides; pronotal carina complete and shortly lamellate, forming a rounded obtuse angle on humeri; pretegular carina sharp; dull oblique humeral carina connecting angle of pronotal carina to ventral extreme of pretegular carina; dorsal faces of pronotum evenly convex, lateral faces depressed ( Fig. 2D ). Mesoscutum as long as wide, lacking notauli or parapsidal sulci, evenly convex in lateral view. Scutellum nearly flat, with weak indication of median furrow near posterior margin; axillary fossa oval, narrow but not slit-like. Metanotum flattened with short lateral carinae. Tegulae with evenly convex outer margin, posterior lobe equaling parategulae and curved down; parategulae evenly curved and apically rounded. Dorsal plate of mesepisternum weakly and evenly convex, ventral plate more strongly convex and with depressed posterior margin; epicnemial carina strong, running from mid coxa to ventral pronotal margin, where it continues in the oblique humeral carina of pronotum. Propodeum evenly and strongly convex in lateral view; dorsal faces convex and smoothly passing into median longitudinal depression; lateral faces flattened; median depression of propodeum with a basal elongate fovea from which a longitudinal carina runs ventrally but not reaching apex of propodeum; lateral carina sharp on posterior half, in dorsal view evenly convex with a barely indicated sinuate lobe above propodeal valvula ( Fig. 2F ). T 1 in dorsal view 3× as long as apically wide, basal three fifths narrow with weakly concave lateral sides, apical two fifths inflated and with straight diverging sides; in lateral view, apical part evenly inflated basally, then abruptly depressed preapically; T1 with a circular transverse carina basally, that curves on sides and continues on lateral faces of tergite and disappear just before apical inflated part, area above the lateral part of carina depressed and giving a “pinched” appearance to the petiole; apical margin forming an upturned brownish lamella that is not distinctly separated from rest of tergite ( Fig. 2G ). S1 narrow and parallel sided basally, then triangularly expanded posteriorly. T2 about as long as wide in dorsal view, with convex sides; strongly and evenly convex in lateral view; apical margin forming a brownish flat lamella separated from rest of tergite by a row of coarse punctures. S2 much less convex than respective tergite.

Clypeus dull on basal half and on sides of apical part, with dense micropunctures and fine barely visible punctures; median depressed area of apical part shiny, with weak irregular microsculpture and scattered indistinct punctures. Frons microreticulate and very dull, with small and shallow but distinct punctures, interspaces width variable from 1× to 4× puncture diameter; vertex and ocular sinus similar to frons, with sparser microreticulation and punctures and a silky shine. Gena similar to vertex on dorsal third, becoming smoother and shinier ventrally. Scape microreticulate and dull, with sparse fine punctures. Mandible dull near base, becoming shiny apically. Dorsal faces of pronotum silky shiny, with weak microreticulation, becoming smooth and shiny just before dorsal carina, punctures large and deep, becoming deeper and denser towards sides but interspaces always flat and at least as wide as puncture diameter; lateral faces of pronotum shiny with weak indication of transverse striae; anterior face of pronotum shiny with fine and sparse micropunctures, some fine punctures along dorsal margin. Mesoscutum, scutellum and metanotum microreticulate with a silky shine, becoming smoother towards posterior margins, punctures fine and deep, denser on anterolateral parts of mesoscutum, sparser and larger on scutellum and metanotum. Mesepisternum shiny with weak microreticulation, sparse fine punctures on dorsal plate, very large foveae present on sulci and along epicnemial carina; epicnemium shiny and micropunctate on sides, reticulate and dull in the middle. Dorsal plate of metaepisternum strongly striate; ventral plate of metaepisternum shiny and microreticulate, with large foveae along anterior margin and some striae along posterior margin. Tegulae microreticulate with scattered fine punctures. Dorsal faces of propodeum microreticulate and subopaque, reticulation becoming progressively coarser from base to apex but disappearing at extreme apex, sparse fine punctures along median and lateral margins; median depression with transverse irregular striae along median carina, apical fifth entirely smooth and shiny; lateral faces of propodeum shiny on dorsal margin and becoming very dull ventrally, covered with fine but distinct transverse striation, with some scattered large shallow punctures. T1 densely microreticulate and dull, reticulation disappearing on inflated part and leaving it smooth, with sparse micropunctation on apical half. T2 dull on basal half, with a silky shine on apical half and shiny on lateral margins, densely microreticulate and micropunctate with scattered deep punctures on sides and along a preapical line separating disc of tergite from the apical lamella, some very shallow and barely visible punctures on disc. T3–T4 sculpted similar to apical part of T2, but micropunctures larger and deeper. T5–T7 weakly reticulate with very sparse fine punctures. S1 with transverse irregular striae along basal narrow part, irregularly rugose and microreticulate on apical triangular part. S2–S6 shiny, reticulation very weak on S2 and becoming stronger towards S6, punctures large and deep on S2 and becoming finer and shallower towards S6. S7 microreticulate and very shiny, with dense fine punctures.

Head and mesosoma with dense silvery setae, longer on frons, metanotum and propodeum, shorter on sides of mesosoma; interantennal area, clypeus and lateral faces of propodeum with short and dense silvery pubescence; metasoma with dense short whitish pubescence, sparser on T1; S1 with long sparse white setae only; T3–T7 and S2–S7 with longer setae mixed in pubescence and denser towards apical margin of sternites.

Apical part of aedeagus subtriangular, more or less evenly narrowing from base to apex, with a rounded apical swell; distinctly curved at apex in lateral view. Ventral lobes of aedeagus in lateral view small and obliquely projecting, apically subtruncate, slightly sinuate distally; subtriangular and laterally projecting in ventral view. Basal plate narrow and evenly tapering apically, pointed at apex. Anteroventral lobe of digitus right angled, apically rounded; distal lobe of digitus subtriangular and evenly tapering to apex; conuli placed in a dense patch along ventral margin of stem of digitus, and in a sparse arched patch on anteroventral lobe. Genitalia in Fig. 2H .

Black; following parts ivory colored: ventral face of scape, irregular spot near base of mandible, subtriangular spot above antennal insertions, small and barely visible spots on basal and apical angles of clypeus, small spot on gena at height of ocular sinus, short transverse lines near middle of pronotal carina, posterior corner of dorsal face of propodeum, small spot on apical corner of T1, very narrow apical band on T2 largely interrupted in the middle and not reaching sides, apical spot on fore femur, line on outer face of fore tibia and basitarsus, irregular and interrupted line on outer face of mid tibia; following parts dark brown-red: mandibles except black margins, labrum, outer margin of tegula, propodeal valvula, apical depressed margins of tergites, most of legs except tarsi becoming lighter towards apex; underside of flagellum orange, darker on F1–F7 and brighter on F8–F11. Wings weakly infuscate along costal margin, venation brown.

Female. Unknown.

Distribution. South Sudan ( Fig. 1 ).

Etymology. The specific name is in reference to the country of origin of this species.

Figures

FIGURE 1. Distributional map of Afrotropical Ectopioglossa species.

FIGURE 2. Ectopioglossa sudanensis Selis, sp. nov., male holotype. A, habitus in dorsal view. B, habitus in lateral view. C, head in frontal view. D, pronotum in dorsolateral view. E, flagellum in lateral view. F, propodeum in dorsolateral view. G, T1 in dorsal view. H, genitalia (aedeagus in ventral view, aedeagus in lateral view, digitus and volsella).