Enaphalodes monzoni, Lingafelter & Santos-Silva, 2018

Lingafelter, Steven W. & Santos-Silva, Antonio, 2018, New Central American and Mexican Enaphalodes Haldeman, 1847 (Coleoptera: Cerambycidae) with taxonomic notes and a key to species, Papéis Avulsos de Zoologia 58, pp. 1-17 : 7-9

Enaphalodes monzoni , sp. nov.

( Figs. 17-23 )

Diagnosis: Enaphalodes monzoni , sp. nov. is very similar to E. bingkirki , sp. nov., due to its short antennae, elytral maculations, and dense, erect, elytral setae. It is distinguished by the antennae just reaching the distal third of the elytra, the prothorax being somewhat wider but without a distinct lateral projection anterior to midpoint ( Fig. 17 ), the pro‑ and mesosternal processes wider ( Fig. 23 ), and the uniformly ochre pubescence dorsally. In E. bingkirki , the antennae reach the distal sixth of the elytra, the prothorax is relatively more slender and with a more pronounced projection laterally just anterior of midpoint ( Fig. 9 ), the pro‑ and mesosternal processes are narrower ( Fig. 24 ), and the dorsal pubescence is a mixture of ochre and yellowish‑white setae. Like E. bingkirki , the dense, erect elytral pubescence in E. monzoni most easily distinguishes it from E.niveitectus .

Description: Holotype female: Integument mostly dark brown, almost black dorsally; mouthparts and legs dark reddish‑brown with darker areas; antennae dark brown onbasal segments,gradually reddish‑browntowarddistal segments. Pubescence and erect setae uniformly ochre.

Head: Frons centrally tumid close to clypeus, with deep fovea on each side close to clypeus; minutely, densely punctate, except nearly smooth tumid area, and narrow band along median groove; pubescence obscuring integument except glabrous tumid area and along median groove, interspersed with a few long, erect setae laterally. Area between antennal tubercles and upper eye lobes with moderately coarse, sparse punctures interspersed with dense, minute punctures; pubescence nearly obscuring integument, except glabrous area along median groove (widened, sub‑diamond‑shaped between upper eye lobes), interspersed with a few long, erect setae close to eyes. Remaining surface of vertex and area behind upper eye lobes, densely, confluently, shallowly punctate; pubescence forming two wide rounded lobes close to eyes, reaching to about middle of area behind upper eye lobes; wide area of vertex and area behind upper eye lobes close to prothorax glabrous. Area behind lower eye lobes striate‑punctate; with band of pubescence close to eye, distinctly widened toward gena. Antennal tubercles flat, rounded at apex; densely, minutely punctate except smooth narrow area close to apex; pubescence nearly obscuring integument, interspersed with sparse, long, erect setae on frontal side, except glabrous smooth area. Genae finely, moderately sparsely punctate; with band of pubescence close to eye, with long, erect, sparse setae on remaining surface. Postclypeus minutely, sparsely punctate centrally, densely punctate at level of fovea on frons, smooth laterally; pubescent close to tumid area on frons, with short and long, moderately sparse setae on remaining surface except glabrous smooth area. Labrum coplanar with anteclypeus on basal ⅔, inclined on distal third; sides of basal ⅔ and entire distal third smooth; remaining surface of basal ⅔ minutely, moderately abundantly punctate close to curvature, smooth basally; with short and long erect setae, denser centrally on inclined area. Gula smooth, glabrous. Submentum vermiculate, interspersed with fine punctures; with moderately sparse decumbent setae interspersed with long, erect setae. Distance between upper eye lobes 0.63 times length of scape; in frontal view, distance between lower eye lobes 0.92 times length of scape. Antennae 1.1 times elytral length, reaching base of distal third of elytra. Scape wide at base and not widened toward apex, flattened and slightly depressed dorsally on basal third, finely, densely punctate; pubescence not obscuring integument, interspersedwith long, erect setae, distinctly more abundant ventrally. Antennomeres dorsoventrally flattened (less so at apex of III‑VI); inner apex of antennomere III with acute spine, slightly narrower than distal width of antennomere; inner apex of antennomere IV with acute spine, with length equal to about 0.75 times distal width of antennomere; apex of antennomere Vwith acute spine, slightly longer than half of distal width of antennomere; antennomeres VI‑X with short spine atinner apex,shorter, gradually shorter toward X; antennomeres V‑X with short spine at outer apex, nearly indistinct on V‑VI; antennomeres with pubescence gradually shorter and denser toward distal segments; antennomeres III‑VII with long, erect setae ventrally (sparser toward VII), and long, erect setae on dorsal and ventral apex of III‑X (gradually shorter toward X). Antennal formula (ratio) basedon length of antennomere III (excluding spine): scape = 1.04; pedicel = 0.16; IV = 0.74; V = 0.90; VI = 0.87; VII = 0.82; VIII = 0.66; IX = 0.64; X = 0.53; XI = 0.64.

Thorax: Prothorax 1.2 times wider than long, with distinct constriction near apex; sides rounded, with small, blunt tubercle at about middle. Pronotum with seven gibbosities: one elongate, placed centrally from near base to apex, distinctly narrowed between middle and distal third; two fused on each side of central gibbosity; one oblique, elongate, irregular, placed on sides of basal half, fused at apex with the latter ones. Pronotal surface coarsely, densely rugose‑punctate, especially laterally, except smooth central gibbosity; pubescence dense on basal sides of central gibbosity, elongate band on out‑ er side of fused gibbosities, strongly curved on top of posterolateral gibbosity (denser on its base and apex), irregular spot on distal area of oblique gibbosity, spot on lateral tubercle, and spot close to anterolateral angle; remaining surface with short, decumbent, sparse setae except glabrous central gibbosity. Sides of prothorax coarsely, densely rugose‑punctate; with short, decumbent, sparse setae, denser closeto prosternum, especially close to procoxal cavity. Prosternum coarsely, densely rugose‑punctate (nearly vermiculate on basal ⅔); with short, sparse, decumbent setae interspersed with long, erect setae, except narrow band with dense pubescence close to procoxal cavities. Prosternal processwith narrow band of densepubescence laterallyon basal half, entirely pubescent on distal half. Mesosternum with pubescence partially obscuring integument. Mesepisternum, mesepimeron and metepisternum with pubescence obscuring integument. Mesosternal process strongly inclined toward mesosternum, with small tubercle on each side closeto curvature. Metasternumcoarsely, moderately abundantly punctate except smooth area close to central sulcus; sides with dense pubescence obscuring integument and wide triangular central area with sparse, erect setae. Scutellum with pubescence obscuring integument.

Elytra: Coarsely, densely punctate on basal third, gradually finer, sparser toward apex; with large, irregular maculae of dense pubescence; with long, erect, sparse setae throughout; apex truncate, bispinose (outer spine slightly longer).

Legs: Femora finely, densely punctate (punctures finer toward apex); pubescence not obscuring integument; apices of profemora with rounded lobe; apices of meso‑ and metafemora with subtriangular lobes.

Abdomen: Ventrites I‑IV with large macula of pubescence on each side of distal half (about distal third on ventrite I); remaining surface with short, sparse, decumbent setae interspersed with long, erect setae (more abundant on V); apex of ventrite V rounded, emarginate centrally.

Dimensions in mm: Total length, 27.15; prothorax: central length, 5.15; distal width, 4.35; basal width, 5.60; widest width, 6.15; humeral width, 8.10; elytral length, 20.00.

Type material: Holotype female from GUATEMALA, Izabal: Sierra del Merendón ( Finca Firmeza , 950 m), 28.VIII‑03.IX.2007, J. Monzón, F. Camposeco and R. E. Woodruff col. ( FSCA) .

Etymology: This species is named after José Monzón who collected the holotype and was host and collaborator to many entomological researchers in Guatemala.

Figures

Figures 17-24. (17‑23): Enaphalodes monzoni, sp.nov., holotype female: (17) Dorsal habitus; (18) Ventral habitus; (19) Lateral habitus; (20) Anteroventral view; (21) Head, frontal view; (22) Ventral side, oblique view; (23) Pro‑ and mesosternal processes. (24) E. bingkirki, sp. nov., holotype female, pro‑ and mesosternal processes.

Figures 9-16. Enaphalodes bingkirki, sp. nov.: (9‑13): Holotype female: (9) Dorsal habitus; (10) Ventral habitus; (11) Lateral habitus; (12) Head, frontal view; (13) Pro‑ and mesosternal process.(14‑16): Paratype female:(14) Lateral habitus;(15) Dorsal habitus;(16)Ventralhabitus.