Epanerchodus bispinosus, Golovatch, Sergei I., Mikhaljova, Elena V. & Chang, Hsueh-Wen, 2011
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https://dx.doi.org/10.3897/zookeys.93.1167 |
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https://treatment.plazi.org/id/1E6074F2-D2D9-0C7A-4A84-5B83F08AF89E |
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scientific name |
Epanerchodus bispinosus |
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sp. n. |
Epanerchodus bispinosus ZBK sp. n. Figs 4553
Type material:
Holotype ♂ (NMNS-6559-001), Taiwan, Nantou County, Ren-ai Township, Mei-Feng, 15.04.2002, leg. S.H. Wu. Paratypes: 3 ♂, 1 ♂ fragm., 4 ♀, 1 ♀ juv. (NMNS-6559-002), 1 ♂, 1 ♀ (ZMUM), 1 ♂ (without head), 1 ♀ (ZMUC), 1 ♂ fragm., 1 ♀ (MNHN JC 334), 1 ♂ (TFRI), 1 ♂ (NSYSU), same locality, together with holotype.
Name:
To emphasize the distal half of the gonopod telopodite being like two long spines.
Diagnosis:
Due to the presence of a strong and spiniform exomere, this new species joins the few congeners hitherto referred to the erstwhile genus Usbekodesmus (see above the synonymy with Epanerchodus ), but differs in the distal half of the gonopod telopodite being represented by only two spiniform branches showing no additional outgrowths (see also Key below).
Description:
Length of both sexes ca 8-11 mm; width of pro- and metazona varying between specimens from 0.8-1.3 to 1.1-1.7 mm, respectively. Holotype ca 9 mm long, and 0.8 and 1.1 mm wide on pro- and metazona, respectively. Coloration in alcohol from pallid to uniformly light grey, yellow or very light red-brown, sometimes head faintly marbled light red-brown; venter and legs yellowish to greyish (Figs 45-49).
All characters as in Epanerchodus orientalis except as follows.
Antennae rather long and evidently clavate, reaching behind end (♂) or midway (♀) of segment 3 dorsally; antennomere 3 longest, considerably longer than a relatively stout, yet highest, 5th (Figs 45); antennomeres 5 and 6 each with a small, compact, distodorsal group of bacilliform sensilla; antennomere 7 with a minute dorsoparabasal cone and a distodorsal group of microscopic sensilla.
In width, collum <segment 2 = 3 <4 <head = 5-15 (Fig. 46), thereafter body gradually tapering towards telson (♂, ♀) (Fig. 48). Paraterga moderately developed, starting from collum, set high but invariably lying slightly below a faintly convex dorsum; paraterga on collum small, subtriangular, with a small lateral incision in front of a rounded caudal corner; front shoulders drawn forward and slightly convex only paraterga 2-4, thereafter straight, increasingly well rounded and directed increasingly caudolaterad; caudal corner on pataterga 2-5 subrectangular and narrowly rounded, starting from 6th increasingly acutangular and beak-shaped, starting from segment 9 first faintly and then extending increasingly beyond rear tergal contour (Figs 45-48). All poreless segments with three, all pore-bearing ones with four, small but evident incisions, each usually bearing a small seta on top at lateral margin. Metatergal sculpture typical, moderately developed, with three indistinct transverse rows of setiferous, polygonal bosses (Figs 46-48). Tergal setae very short, mostly retained, a little longer only on collum and in rear row on metatergum 19. Stricture between pro- and metazona wide and smooth. Limbus very thin, microdenticulate. Epiproct rather short, conical (Fig. 48), only slightly bent ventrad, preapical papillae evident. Hypoproct semi-circular; caudal, paramedian, setiferous papillae evident and well-separated.
Sterna without modifications, very densely setose. Legs rather short, slender, in ♂ evidently enlarged (Figs 45), ca 1.5-1.6 (♂) or 1.1-1.2 times (♀) as long as midbody height, ♂ prefemora not swollen dorsally, together with femora beset with bifid setae ventrally, acropodite (postfemur+tibia+tarsus) with sphaerotrichomes ventrally (Fig. 51).
Gonopods (Figs 50, 52, 53) with large, subquadrate, medially fused coxae carrying only a few long setae ventrally. Telopodite slender and simple, prefemoral portion relatively short, exomere (ex) spiniform, endomere (en) only a little longer than ex, apex bent unciform mesally, devoid of any outgrowths near base; hairy pulvillus evident.
Remarks.
This species is remarkable in showing an exomere, albeit usual and spiniform, almost as long as a particularly simple endomere. This condition nicely bridges the weak distinction which has hitherto remained to formally keep Usbekodesmus and Epanerchodus as independent genera ( Geoffroy and Golovatch 2004), with the gonopod structure in Epanerchodus bispinosus sp. n. providing the final evidence to formally synonymize these genera (see also above).
Epanerchodus bispinosus sp. n. in Taiwan is apparently very local in distribution, having been encountered only at a single locality (Map 2).
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.
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