Estheria pallicornis ( Loew, 1873 )

Zhang, Chun-Tian, Shima, Hiroshi, Liang, Hou-Can & Li, He-Nan, 2019, Four new species of Estheria Robineau-Desvoidy (Diptera: Tachinidae) from China and Nepal, with a review of the East Palearctic and Oriental species, Zootaxa 4603 (1), pp. 1-38 : 21-23

Estheria pallicornis ( Loew, 1873)

( Figs 11 , 23 , 35 , 47 , 63 , 72 , 89 , 102 , 117 )

Dinera pallicornis Loew, 1873: 237 . Type locality: Uzbekistan or Tajikistan, Zarafshon Valley [as “ Sarawschan-Thal ”]. Type (s), male (1 in ZMHB, J. Ziegler, pers. comm.), not examined. References: Kolomiets (1973: 98; key and description); Crosskey (1976: 180; Oriental catalog); Herting & Dely-Draskovits (1993: 364; Palearctic catalog); Cerretti & Tschorsnig (2012: 273, 284; species and key); Zhang et al. (2013: 139; checklist).

Dolichodexia rufipes Brauer & Bergenstamm, 1889: 118 and 167 (key). Type locality: Turkey, Amasya.

Dolichodexia albipila Mesnil, 1963: 54 . Type locality: Tajikstan, Khorog. Reference: Kolomiets (1973: 97).

Diagnosis. Prementum short; antennae, palpi and legs reddish-yellow; pleura, postpronotal lobes, coxae and base of abdomen ventrally with dense pale hairs; veins M and R 4+5 merged to form a short petiole.

Redescription. Body length: 8.8–14.3 mm.

Male. Head covered with grayish-white pruinosity, frontal vitta dark brown, lunule and arista brown; antenna, palpus, prementum and labella reddish-yellow. Frons 0.1–0.12 of head width, widened anteriorly; frontal vitta narrower than fronto-orbital plate at middle, latter with some fine black hairs; parafacial bare, 3–3.5 times as wide as 1 st flagellomere ; facial carina low and indistinct; gena as long as palpus or antenna, 0.42–0.57 of eye height; lower margin of face not protruding forward. Fronto-orbital plate with short pale and dark hairs, parafacial bare; inner vertical seta 0.3–0.4 of eye height, shorter than ocellar seta; outer vertical seta hair-like; 10–11 frontal setae; orbital seta absent; ocellar seta longer than upper frontal seta; vibrissa inserted above lower margin of face; a row of subvibrissa below vibrissa; upper occiput without black hairs behind postocular setae. Antenna falling short of lower margin of face by more than length of antenna, 1 st flagellomere 1.5–2 times as long as pedicel; arista, including plumosity, 3–4 times as wide as first flagellomere; palpus about as long as 1 st flagellomere ; prementum short, about twice as long as wide or as long as palpus.

Thorax brown to dark brown, with yellowish-gray pruinosity; scutellum dark brown, with gray-white pruinosity; postprontal lobe dark brown; pleura with gray pruinosity; anterior and posterior spiracles reddishyellow. Dorsum with 4 longitudinal black vittae extending back to anterior half of postsutural scutum, narrower inner vitta 1/3–1/4 width of pruinose portion between inner and outer vittae; dorsum with black hairs except for pale white hairs on anterior and outer-lateral postprontal lobe; pleura with dense pale hairs, without black hairs. Prosternum bare, about 1.5 times as long as wide; proepisternum bare; postpronotal lobe with 4–5 setae, 3 of which nearly in a straight line; 2–3 presutural and 3 postsutural acrostichal setae; 3 presutural and 3 postsutural dorsocentral setae; 3 postsutural intra-alar setae; prealar setae weak, 2 supra-alar setae [1 male from Pakistan with 3–5 supra-alar setae]; 3 katepisternal setae; katepimeron hairy; posterior lappet of posterior spiracle circular; 3 pairs of marginal scutellar setae and 1 pair of discal scutellar setae. Wing hyaline, brownish; tegula and basicosta dark brown; halter reddish-yellow. Calypters yellowish-white; lower calypter with short fringe on outer margin. Costal spine absent; second costal sector bare ventrally, relative length of 2nd, 3rd and 4th costal sectors approximately 1.2:2.5:1; bend of vein M distinctly more than right-angled [specimens from China] or nearly rightangled [specimens from Pakistan]; vein M from crossvein dM-Cu to bend about 3/4 times as long as distance between bend and wing margin, with an appendage longer than crossvein r-m; vein M merged with vein R 4+5 to form a short petiole. Legs with femora brown-yellow except for ventral apex with grayish-white pruinosity, tibiae reddish-yellow, tarsi dark brown, pulvilli yellowish; coxae and ventral surface of femora with pale yellowish hairs. Fore tibia with 3–5 very short anterodorsal setae on basal 1/2 and 2 median posterior setae; fore claws and pulvilli slightly longer than 5th tarsomere; mid tibia with 1 anteroventral, 1 anterodorsal and 2 posterodorsal setae; hind tibia with 2 anteroventral, 3–4 anterodorsal and 2–3 posterodorsal setae, 1 apical anteroventral and 2 apical dorsal setae.

Abdomen long-ovate, black, covered with yellowish-gray pruinosity, dark brown on posterior margin around setae, with pale long hairs on ventral surface at base; syntergite 1+2 medially excavated to posterior margin, with 1 lateral marginal seta, without median marginal setae; tergite 3 with 2 median marginal setae, 1–2 lateral marginal setae, and 2 median discal setae; tergite 4 with a row of marginal setae and 2 median discal setae; tergite 5 with a row of marginal setae and many discal setae. Sternites with erect pale white hairs. Male terminalia ( Figs. 11 , 23 , 35 , 47 ). Posterior lobe of sternite 5 wide and bluntly rounded, V-shaped median cleft deep and wide. Surstylus short, bluntly round at apex; cerci separated, long, narrow and pointed apically; basiphallus about as long as postgonite; distiphallus long and slender, membranous apical part about 1/3 as long as sclerotized basal part; ejaculatory apodeme small.

Female. As male, except as follows: postprontal lobe reddish brown, scutellum brown, with grayish-white pruinosity; legs yellow including coxae and tarsi; pulvilli yellowish, claws reddish-yellow on basal half and brown to dark in apical half; abdomen with grayish-white pruinosity, tergites 3 to 5 black along posterior margin. Frons 0.38–0.39 of head width; frontal vitta about as wide as fronto-orbital plate; parafacial about 4 times as wide as 1st flagellomere; gena 0.42–0.57 of eye height; inner vertical seta stronger than ocellar setae; outer vertical seta about 3/5 as long as inner vertical seta; 7 inclinate frontal setae, about as long as ocellar setae; 2 strong proclinate orbital setae and 1 lateroclinate prevertical seta. Fore claws and pulvilli shorter than 5th tarsomere; mid tibia with 1–2 anterodorsal setae; hind tibia with 2–4 anterodorsal setae. Abdomen ovate, without median discal setae on tergites 3 to 5.

Material examined. China, Beijing: 2 ♂♂, Mt. Baihua , 1170–1520 m, 24.VIII.1963, S.-Y. Wang leg. ( IZCAS) ; 2 ♂♂, Wofosi , 5.X.1971, J.-M. Chao leg. ( IZCAS) ; 1 ♂, Badaling , 3.IX.1980, E.-Y. Liang leg. ( IZCAS) ; 12 ♂♂, 32 ♀♀, Xiatougou , 6–12.X.1982, Z.-C. Su leg. ( IZCAS) ; 1 ♂, 1 ♀, Mentougou , 4–6.IX.1976, F.-S. Li leg. ( CAU) ; 1 ♀, Mentougou , 8.VIII.2008, T. Wang leg. ( CAU) . Hebei: 2 ♂♂, Xiaowutai Mountains, Beitai-Tatou , 1200–2400 m, 14–22.VIII.1964, C. Wang & Y.-H. Han leg. ( IZCAS) ; 1 ♂, same data as previous except VIII.1980 ( IZCAS) ; 1 ♂, same data as previous except 26.VI.2009, C. Fu leg. ( SYNU) . Inner Mongolia: 1 ♂, Jining , Slope, Hardwood, 1.VIII.1971, Y.-S. Shi leg. ( IZCAS) ; 1 ♂, Xi Ujimqin Qi , Grassland, 6.VIII.1971, Y.-S. Shi leg. ( IZCAS) ; 1 ♀, Tumuertai , 12.VIII.1979, Y.-S. Shi leg. ( IZCAS) ; 1 ♀, Zhungeer , 18.VIII.2008, F.-Y. Yang leg. ( IZCAS) ; 4 ♂♂, 4 ♀♀, Alashanzuo Qi, Helan Mountains, Halawu , 2200–2800 m, 25.VII–6.VIII.2010, S.-D. Wang & Z. Zhao leg. ( SYNU) ; 5 ♂♂, 3 ♀♀, Helanshan, Xiangchizi , 2000–2500 m, 6–7.VIII.2011, H. Lin & C. Tan leg. ( SYNU) ; 7 ♂♂, 2 ♀♀, Baotou, Shala Qi, Goumen , 17.VIII.2015 and 2.?.2015, A.-P. Liu leg. ( IGRCAAS) ; 2 ♂♂, 1 ♀, Hohhot, Wuchuan, Daqingshan , Shierdeng Village , 16–21.?.2017, D.-H. Wang leg. ( IMAU) . Shanxi: 4 ♀♀, Pinglu, Xiashuitou , 2.IX.1979, M.-F. Wang leg. ( SYNU) ; 11 ♂♂, 3 ♀♀, Guangling, Shangbaiyang , 9.VIII.1980, M.-F. Wang leg. ( SYNU) ; 8 ♂♂, Shanyin, Xialakou , 11.VII.1982, M.-F. Wang leg. ( SYNU) ; 11 ♂♂, Yangquan, Shanyin, Shangqiu , 12.VII.1982, M.-F. Wang leg. ( SYNU) ; 1 ♂, Guangling, Xianglutai , 19.VII.1985, M.-F. Wang leg. ( SYNU) ; 1 ♂, Zuoquan , 12.VI.1999, M.-F. Wang leg. ( SYNU) . Ningxia: 1 ♂, Jingyuan, Helan Mountains, Xixia Forestry Center , 2000 m, 11.VII.2008, Z.-Y. Yao leg. ( SYNU) . Xinjiang: 1 ♂, Zhaosu , 1630 m, 11.VIII.1957, H. Wang leg. ( IZCAS) ; 2 ♂♂, same data as previous except 15.VI.1959, S.-Y. Wang leg. ( IZCAS) ; 1 ♀, Burjin , 480 m, 18.VII.1960, S.-Y. Wang leg. ( IZCAS) ; 2 ♂♂, 7 ♀♀, Manasi , 400–900 m, 9.VI.1963, S.-Y. Wang leg. ( IZCAS) ; 1 ♀, Manasi , 9.VII.1967, C.-P. Hong leg. ( IZCAS) ; 1 ♂, Zhaosu, Xiata , 1950 m, 18.VII.1978, X.-Z. Zhang leg. ( IZCAS) ; 1 ♂, Hanasi Lake , 15.VII.1980, X.-W. Liu leg. ( SEMCAS) ; 24 ♂♂, 3 ♀♀, Shule , 1280 m, 10–22.V.2007, M.-L. Sheng leg. ( GSFPM) ; 1 ♀, Nalati , 6.VIII.2007, S. Huo leg. ( GSFPM) . Nepal: 1 ♀, Phulchouki , 2500 m, 1.XI.1992, M. Iwasa leg. ( BLKU) . Pakistan: 1 ♂, Swat Ushu , 2000 m, 10 km N of Kalam, 10.VIII.1987, H. Kurahashi leg. ( BLKU) ; 1 ♀, Kalan , 90 km N of Mingola, 10.VIII.1987, K. Kanmiya leg. ( BLKU) ; 1 ♀, 6 km N of Gilgit , 27.VII.1987, K. Kanmiya leg. ( BLKU) ; 1 ♀, Ultar gracier, Hunza , 2500 m, 31.VII.1987, S. Shinonaga leg. ( BLKU) ; 1 ♀, Punjab, Fort Munro , 3.IX.1988, M. Iwasa leg. ( BLKU) ; 2 ♀♀, Baluchistan, Ziarat , 17.IX.1988, M. Iwasa leg. ( BLKU) ; 2 ♀♀, NWF Prov., Hopel nr Gakuch , 1900 m, 6.VII.1989, H. Kurahashi leg. ( BLKU) .

Distribution. Afghanistan, China (Beijing, Hebei, Inner Mongolia, Ningxia, Shanxi, Xinjiang), Greece, India, Iran, Kyrgyzstan, Mongolia, Nepal *, Pakistan, Russia (W and E Siberia), Tajikstan, Transcaucasia, Turkey, Uzbekstan.

Hosts. Scarabaeidae : Amphimallon solstitiale (Linnaeus) , Brahmina agnella (Faldermann) , Lasiopsis henningi (Fischer) , Melolontha hippocastani Fabricius and Mimela holosericea (Fabricius) ( Tschorsnig 2017: 342) .

Remarks. This species is easily distinguished from other species of this genus in having prementum short, antenna, palpi and legs reddish-yellow, long and dense pale hairs on pleura, postpronotal lobe, coxae and ventral base of abdomen.

Figures

FIGURES 1–12. Male terminalia of Estheria spp. in caudal view. 1. E. hirtinerva sp. nov., paratype from Geka-Echu Mountains, Daocheng, Sichuan, China (SYNU). 2. E. prostata sp. nov., paratype from Gezan, Shangri-La, Yunnan, China (SYNU). 3. E. tibetensis sp. nov., paratype from Geka- Echu Mountains, Daocheng, Sichuan, China (SYNU). 4. E. wangi sp. nov., paratype from Napahai wetland, Shangri-La, Yunnan, China (SYNU). 5. E. acuta (Portschinsky), specimen from Hunyuan, Shanxi, China (SYNU). 6. E. cristata (Meigen), specimen from Baden-Willingen, Germany (SMNS). 7. E. decolor (Pandellé), specimen from Dongsheng, Inner Mongolia, China (SYNU). 8. E. flavipennis Herting, specimen from Xi Ujimqin Qi, Inner Mongolia, China (IZCAS). 9. E. maculipennis Herting, specimen from Dong Ujimqin Qi, Inner Mongolia, China (IZCAS). 10. E. magna (Baranov), specimen from Mt. Fenghuang, Fengcheng, Liaoning, China (SYNU). 11. E. pallicornis (Loew), specimen from Shule, Xinjiang, China (GSFPM). 12. E. petiolata (Bonsdorff), specimen from Kanas Lake, Burqin, Xinjiang, China (SEMCAS). Scale bars = 0.2 mm.

FIGURES 13–24. Male terminalia of Estheria spp. in lateral view. 13. E. hirtinerva sp. nov., paratype from Geka-Echu Mountains, Daocheng, Sichuan, China (SYNU). 14. E. prostata sp. nov., paratype from Gezan, Shangri-La, Yunnan, China (SYNU). 15. E. tibetensis sp. nov., paratype from Geka- Echu Mountains, Daocheng, Sichuan, China (SYNU). 16. E. wangi sp. nov., paratype from Napahai wetland, Shangri-La, Yunnan, China (SYNU). 17. E. acuta (Portschinsky), specimen from Hunyuan, Shanxi, China (SYNU). 18. E. cristata (Meigen), specimen from Baden-Willingen, Germany (SMNS). 19. E. decolor (Pandellé), specimen from Dongsheng, Inner Mongolia, China (SYNU). 20. E. flavipennis Herting, specimen from Xi Ujimqin Qi, Inner Mongolia, China (IZCAS). 21. E. maculipennis Herting, specimen from Dong Ujimqin Qi, Inner Mongolia, China (IZCAS). 22. E. magna (Baranov), specimen from Mt. Fenghuang, Fengcheng, Liaoning, China (SYNU). 23. E. pallicornis (Loew), specimen from Shule, Xinjiang, China (GSFPM). 24. E. petiolata (Bonsdorff), specimen from Kanas Lake, Burqin, Xinjiang, China (SEMCAS). Scale bars = 0.2 mm.

FIGURES 25–36. Estheria spp., phallic complexes in lateral view. 25. E. hirtinerva sp. nov., paratype from Geka-Echu Mountains, Daocheng, Sichuan, China (SYNU). 26. E. prostata sp. nov., paratype from Gezan, Shangri-La, Yunnan, China (SYNU). 27. E. tibetensis sp. nov., paratype from Geka- Echu Mountains, Daocheng, Sichuan, China (SYNU). 28. E. wangi sp. nov., paratype from Napahai wetland, Shangri-La, Yunnan, China (SYNU). 29. E. acuta (Portschinsky), speciumen from Hunyuan, Shanxi, China (SYNU). 30. E. cristata (Meigen), specimen from Baden-Willingen, Germany (SMNS). 31. E. decolor (Pandellé), specimen from Dongsheng, Inner Mongolia, China (SYNU). 32. E. flavipennis Herting, specimen from Xi Ujimqin Qi, Inner Mongolia, China (IZCAS). 33. E. maculipennis Herting, specimen from Dong Ujimqin Qi, Inner Mongolia, China (IZCAS). 34. E. magna (Baranov), specimen from Mt. Fenghuang, Fengcheng, Liaoning, China (SYNU). 35. E. pallicornis (Loew), specimen from Shule, Xinjiang, China (GSFPM). 36. E. petiolata (Bonsdorff), specimen from Kanas Lake, Burqin, Xinjiang, China (SEMCAS). Scale bars = 0.2 mm.

FIGURES 37–52. Male terminalia of Estheria spp. 37–48, 52. Sternite 5 in ventral view. 49–50. Epandrial complex in caudal (49) and lateral (50) views. 51. Phallic complex in lateral view. 37. E. hirtinerva sp. nov., paratype from Geka-Echu Mountains, Daocheng, Sichuan, China (SYNU). 38. E. prostata sp. nov., paratype from Gezan, Shangri-La, Yunnan, China (SYNU). 39. E. tibetensis sp. nov., paratype from Geka- Echu Mountains, Daocheng, Sichuan, China (SYNU). 40. E. wangi sp. nov., paratype from Napahai wetland, Shangri-La, Yunnan, China (SYNU). 41. E. acuta (Portschinsky), specimen from Hunyuan, Shanxi, China (SYNU); 42. E. cristata (Meigen), specimen from Baden-Willingen, Germany (SMNS). 43. E. decolor (Pandellé), specimen from Dongsheng, Inner Mongolia, China (SYNU). 44. E. flavipennis Herting, specimen from Xi Ujimqin Qi, Inner Mongolia, China (IZCAS). 45. E. maculipennis Herting, specimen from Dong Ujimqin Qi, Inner Mongolia, China (IZCAS). 46. E. magna (Baranov), specimen from Mt. Fenghuang, Fengcheng, Liaoning, China (SYNU). 47. E. pallicornis (Loew), specimen from Shule, Xinjiang, China (GSFPM). 48. E. petiolata (Bonsdorff), specimen from Kanas Lake, Burqin, Xinjiang, China (SEMCAS). 49–52. E. picta (Meigen), specimen from Dong Ujimqin Qi, Inner Mongolia, China (IZCAS). Scale bars = 0.2 mm.

FIGURES 53–64. Estheria spp., male heads in anterior view. 53. E. hirtinerva sp. nov., paratype from Solukhumbu District, Nepal (BLKU). 54. E. prostata sp. nov., holotype (SYNU). 55. E. tibetensis sp. nov., paratype from Pianma, Yakou, Lushui, Yunnan, China (SYNU). 56. E. wangi sp. nov., holotype (SYNU). 57. E. acuta (Portschinsky), specimen from Hunyuan, Shanxi, China (SYNU). 58. E. cristata (Meigen), specimen from Baden-Willingen, Germany (SMNS). 59. E. decolor (Pandellé), specimen from Dongsheng, Inner Mongolia, China (SYNU). 60. E. flavipennis Herting, specimen from Xi Ujimqin Qi, Inner Mongolia, China (IZCAS). 61. E. maculipennis Herting, specimen from Dong Ujimqin Qi, Inner Mongolia, China (IZCAS). 62. E. magna (Baranov), specimen from Baishilazi, Kuandian, Liaoning, China (SYNU). 63. E. pallicornis (Loew), specimen from Shule, Xinjiang, China (GSFPM). 64. E. petiolata (Bonsdorff), specimen from Briançon, Nevache, France (SMNS).

FIGURES 65–73. Estheria spp., female heads in anterior view. 65. E. hirtinerva sp. nov., paratype from Duoxiongla Mountain, Paizhen, Milin to Médog, Tibet, China (SYNU). 66. E. prostata sp. nov., paratype from Xuekagou, Milin, Tibet, China (SYNU). 67. E. tibetensis sp. nov., paratype from Pianma, Yakou, Lushui, Yunnan, China (SYNU). 68. E. wangi sp. nov., paratype from Xuekagou, Milin, Tibet, China (SYNU). 69. E. cristata (Meigen), specimen from Baden-Willingen, Germany (SMNS). 70. E. flavipennis Herting, specimen from Huanren, Laotudingzi Mountains, Liaoning, China (SYNU). 71. E. magna (Baranov), specimen from Baishilazi, Kuandian, Liaoning, China (SYNU). 72. E. pallicornis (Loew), specimen from Alashanzuo Qi, Helan Mountain, Halawu, Inner Mongolia, China (SYNU). 73. E. petiolata (Bonsdorff), specimen from Briançon, Nevache, France (SMNS). Scale bars = 0.5 mm (Figs 65–66, 69–72); 1.0 mm (Figs 67–68, 73).

FIGURES 83–91. Estheria spp., male heads in lateral view. 83. E. acuta (Portschinsky), specimen from Hunyuan, Shanxi, China (SYNU). 84. E. cristata (Meigen), specimen from Baden-Willingen, Germany (SMNS). 85. E. decolor (Pandellé), specimen from Dongsheng, Inner Mongolia, China (SYNU). 86. E. flavipennis Herting, specimen from Xi Ujimqin Qi, Inner Mongolia, China (IZCAS). 87. E. maculipennis Herting, specimen from Dong Ujimqin Qi, Inner Mongolia, China (IZCAS). 88. E. magna (Baranov), specimen from Baishilazi, Kuandian, Liaoning, China (SYNU). 89. E. pallicornis (Loew), specimen from Alashanzuo Qi, Helan Mountain, Halawu, Inner Mongolia, China (SYNU). 90. E. petiolata (Bonsdorff), specimen from Briançon, Nevache, France (SMNS). 91. E. picta (Meigen), specimen from Dong Ujimqin Qi, Inner Mongolia, China (IZCAS). Scale bars: 0.5 mm (Figs 87–89); 1 mm (Figs 90–91).

FIGURES 92–106. Estheria spp., male habitus in dorsal (92–104) and lateral (106) view; head in anterior view (105). 92. E. hirtinerva sp. nov., paratype from Solukhumbu District, Nepal (BLKU). 93. E. prostata sp. nov., holotype (SYNU). 94. E. tibetensis sp. nov., paratype from Pianma, Yakou, Lushui, Yunnan, China (SYNU). 95. E. wangi sp. nov., holotype (SYNU). 96. E. acuta (Portschinsky), specimen from Hunyuan, Shanxi, China (SYNU). 97. E. cristata (Meigen), specimen from Baden- Willingen, Germany (SMNS). 98. E. decolor (Pandellé), specimen from Dongsheng, Inner Mongolia, China (SYNU). 99. E. flavipennis Herting, specimen from Xi Ujimqin Qi, Inner Mongolia, China (IZCAS). 100. E. maculipennis Herting, specimen from Dong Ujimqin Qi, Inner Mongolia, China (IZCAS). 101. E. magna (Baranov), specimen from Baishilazi, Kuandian, Liaoning, China (SYNU). 102. E. pallicornis (Loew), specimen from Alashanzuo Qi, Helan Mountain, Halawu, Inner Mongolia, China (SYNU). 103. E. petiolata (Bonsdorff), specimen from Briançon, Nevache, France (SMNS). 104–106. E. picta (Meigen), specimen from Dong Ujimqin Qi, Inner Mongolia, China (IZCAS).

FIGURES 107–118. Estheria spp., male habitus in lateral view. 107. E. hirtinerva sp. nov., paratype from Solukhumbu District, Nepal (BLKU). 108. E. prostata sp. nov., holotype (SYNU). 109. E. tibetensis sp. nov., paratype from Pianma, Yakou, Lushui, Yunnan, China (SYNU). 110. E. wangi sp. nov., holotype (SYNU). 111. E. acuta (Portschinsky), specimen from Hunyuan, Shanxi, China (SYNU). 112. E. cristata (Meigen), specimen from Baden-Willingen, Germany (SMNS). 113. E. decolor (Pandellé), specimen from Dongsheng, Inner Mongolia, China (SYNU). 114. E. flavipennis Herting, specimen from Xi Ujimqin Qi, Inner Mongolia, China (IZCAS). 115. E. maculipennis Herting, specimen from Dong Ujimqin Qi, Inner Mongolia, China (IZCAS). 116. E. magna (Baranov), specimen from Baishilazi, Kuandian, Liaoning, China (SYNU). 117. E. pallicornis (Loew), specimen from Alashanzuo Qi, Helan Mountain, Halawu, Inner Mongolia, China (SYNU). 118. E. petiolata (Bonsdorff), specimen from Briançon, Nevache, France (SMNS).