Euphorbia splendens Bojer ex Hook. var. breonii Leandri

Haevermans, Thomas, Rouhan, Germinal, Hetterscheid, Wilbert, Teissier, Marc, Belarbi, Karim, Aubriot, Xavier & Labat, Jean-Noël, 2009, Chaos revisited: nomenclature and typification of the Malagasy endemic Euphorbia subgenus Lacanthis (Raf.) M. G. Gilbert, Adansonia (3) 31 (2), pp. 279-299 : 295-298

126. Euphorbia splendens Bojer ex Hook. var. breonii Leandri

Notulae Systematicae 12: 159 (1946), nomen, and Cactus (Paris) 34: 114 (1952). — Euphorbia milii Des Moul.var. breonii Ursch & Leandri , nom. nud., Mémoires de l’Institut scientifique de Madagascar, série B, Biologie végétale 5: 148, fig. 38 (1954).

REMARK

According to Newton (1994), Euphorbia breonii Nois. is a superfluous renaming of E. milii Des Moul. , but the epithet was misapplied to a new species by Leandri (1946: 159), without validly describing the taxon (see the section “Names invalidly published by Leandri (1946) and Ursch & Leandri (1954) ”). Consequently, Euphorbia splendens Bojer ex Hook. var. breoni and Euphorbia milii Des Moul.var. breonii were invalidly published nomina nuda, excluding the type of their epithet-bringing synonym.

NAMES INVALIDLY PUBLISHED

BY LEANDRI 1946

AND URSCH & LEANDRI 1954

Leandri (1946) introduced in an identification key,

four new varieties of Euphorbia splendens Bojer ex

Hook. without descriptions. In their work on the

Euphorbia from the botanical garden of Tsimbazaza,

Ursch & Leandri (1954) revised this group. They

based new combinations on these three names but,

this time again, did not give any Latin descriptions.

These names are not validly published because they

were not and still have not been “accompanied by

a Latin description or diagnosis or by a reference to

a (…) Latin description or diagnosis” ( McNeill et

al. 2006: Art. 36):

Euphorbia splendens Bojer ex Hook. var. betsileana Leandri, Notulae Systematicae 12: 159 (1946).

Euphorbia splendens Bojer ex Hook. var. breoni Leandri, Notulae Systematicae 12: 159 (1946).

Euphorbia splendens Bojer ex Hook. var. imperatae Leandri, Notulae Systematicae 12: 159 (1946).

Euphorbia splendens Bojer ex Hook. var. tananarivae Leandri, Notulae Systematicae 12: 159 (1946).

Euphorbia splendens Bojer ex Hook. var. vulcani Leandri, Notulae Systematicae 12: 159 (1946).

Euphorbia milii Des Moul. var. breonii Ursch & Leandri, Mémoires de l’Institut scientifique de Madagascar, série B, Biologie végétale 5: 148, fig. 38 (1954).

Euphorbia milii Des Moul. var. imperatae (Leandri) Ursch & Leandri, Mémoires de l’Institut scientifique de Madagascar, série B, Biologie végétale 5: 150 (1954).

Euphorbia milii Des Moul. var. tananarivae (Leandri) Ursch & Leandri, Mémoires de l’Institut scientifique de Madagascar, série B, Biologie végétale 5: 150 (1954).

Euphorbia milii Des Moul. var. vulcani (Leandri) Ursch & Leandri, Mémoires de l’Institut scientifique de Madagascar, série B, Biologie végétale 5: 150 (1954).

NAMES INVALIDLY PUBLISHED

BY W. RAUH IN SCHILL 1971

In a cytotaxonomic study of the genus Euphorbia L., Schill (1971) mentions two varieties of spiny euphorbias, announcing their description by Rauh in preparation. These species have never been published and their names are therefore two unpublished nomina nuda:

Euphorbia decaryi var. tulearensis Rauh , nomen nudum Euphorbia primulifolia var. isaloensis Rauh , nomen

nudum (as “ primulaefolia ”).

INDEX TO VALID AND INVALID NAMES ACCEPTED NAMES IN BOLD

Euphorbia alfredii Rauh

Euphorbia ambarivatoensis Rauh & Bard. -Vauc.

Euphorbia ambovombensis Rauh & Razaf. var. ambovombensis

Euphorbia ambovombensis Rauh & Razaf. var. ambatomenaensis Rebmann

Euphorbia analavelonensis Rauh & Mangelsdorff

Euphorbia ankarensis Boiteau

Euphorbia ankazobensis Rauh & Hofstätter

Euphorbia annamarieae Rauh

Euphorbia antankara Leandri = Euphorbia pachypodioides Boiteau

Euphorbia aureoviridiflora (Rauh) Rauh

Euphorbia bailloni Boiss. mss. = Euphorbia boissieri Baill.

Euphorbia banae Rauh

Euphorbia beharensis Leandri var. adpressifolia Rauh

Euphorbia beharensis Leandri var. beharensis

Euphorbia beharensis Leandri var. guillemetii (Ursch & Leandri) Rauh

Euphorbia beharensis Leandri var. squarrosa Rauh

Euphorbia beharensis Leandri var. truncata Rauh

Euphorbia berevoensis Lawant & Buddens.

Euphorbia berorohae Rauh & Hofstätter

Euphorbia bevilaniensis Croizat = Euphorbia milii Des Moul. var. bevilaniensis (Croizat) Ursch & Leandri

Euphorbia biaculeata Denis

Euphorbia boissieri Baill.

Euphorbia boiteaui Leandri

Euphorbia bojeri Hook. = Euphorbia milii Des Moul. var. milii

Euphorbia brachyphylla Denis

Euphorbia breoni Nois. = Euphorbia milii Des Moul. var. milii

Euphorbia bulbispina Rauh & Razaf.

Euphorbia capmanambatoensis Rauh

Euphorbia capsaintemariensis Rauh var. capsaintemariensis = Euphorbia decaryi var. capsaintemariensis (Rauh) Cremers

Euphorbia capsaintemariensis Rauh var. tulearensis Rauh = Euphorbia tulearensis (Rauh) Rauh

Euphorbia capuronii Ursch & Leandri

Euphorbia caput-aureum Denis

Euphorbia cremersii Rauh & Razaf. var. cremersii

Euphorbia cremersii Rauh & Razaf. forma viridiflora Rauh

Euphorbia cremersii Rauh & Razaf. var. rakotozafyi (Cremers) Rauh

Euphorbia croizatii Leandri

Euphorbia cylindrifolia Marn. -Lap. & Rauh subsp. cylindrifolia

Euphorbia cylindrifolia Marn. -Lap. & Rauh subsp. tuberifera Rauh

Euphorbia daphnoides Baill. = Euphorbia hexadenia Denis

Euphorbia decaryi Guillaumin var. ampanihyensis Cremers

Euphorbia decaryi Guillaumin var. capsaintemariensis (Rauh) Cremers

Euphorbia decaryi Guillaumin var. decaryi

Euphorbia decaryi Guillaumin var. robinsonii Cremers

Euphorbia decaryi Guillaumin var. spirosticha Rauh & Buchloh

Euphorbia decaryi Guillaumin var. tulearensis Rauh

Euphorbia delphinensis Ursch & Leandri

Euphorbia denisiana Guillaumin

Euphorbia didiereoides Denis ex Leandri

Euphorbia duranii Ursch & Leandri var. ankaratrae Ursch & Leandri

Euphorbia duranii Ursch & Leandri var. duranii

Euphorbia erythrocucullata Mangelsdorff

Euphorbia fianarantsoae Ursch & Leandri

Euphorbia fournieri Rebut = Euphorbia leuconeura Boiss.

Euphorbia francoisii Leandri var. crassicaulis Rauh

Euphorbia francoisii Leandri var. crassicaulis forma rubrifolia Rauh

Euphorbia francoisii Leandri var. francoisii

Euphorbia francoisii Leandri var. rakotozafyi Cremers = Euphorbia cremersii Rauh & Razaf. var. rakotozafyi (Cremers) Rauh

Euphorbia genoudiana Ursch & Leandri

Euphorbia geroldii Rauh

Euphorbia gottlebei Rauh

Euphorbia guillauminiana Boiteau

Euphorbia guillemetii Ursch & Leandri = Euphorbia beharensis Leandri var. guillemetii (Ursch & Leandri) Rauh

Euphorbia hermanschwartzii Rauh

Euphorbia hexadenia Denis

Euphorbia hislopii N.E.Br. = Euphorbia milii Des Moul. var. hislopii (N.E.Br.) Ursch & Leandri

Euphorbia hofstaetteri Rauh

Euphorbia horombensis Ursch & Leandri

Euphorbia iharanae Rauh

Euphorbia isalensis Leandri

Euphorbia isaloensis Drake

Euphorbia itremensis Kimnach & Lavranos

Euphorbia kondoi Rauh & Razaf.

Euphorbia labatii Rauh & Bard. -Vauc.

Euphorbia leandriana Boiteau

Euphorbia leuconeura Boiss.

Euphorbia lophogona Hort. = Euphorbia leuconeura Boiss.

Euphorbia lophogona Lam. var. lophogona

Euphorbia lophogona Lam. var. tenuicaulis Rauh

Euphorbia mahafalensis Denis var. mahafalensis

Euphorbia mahafalensis Denis var. xanthadenia (Denis) Leandri

Euphorbia mainiana Poiss.

Euphorbia mangelsdorffii Rauh

Euphorbia mangokyensis Denis

Euphorbia melanacantha Drake = Euphorbia milii Des Moul. var. splendens (Bojer ex Hook.) Ursch & Leandri

Euphorbia milii Des Moul. var. bosseri Rauh = Euphorbia neobosseri (Rauh) Rauh

Euphorbia milii Des Moul. var. breonii Ursch & Leandri

Euphorbia milii Des Moul. var. imperatae (Leandri) Ursch & Leandri

Euphorbia milii Des Moul. var. longifolia Rauh

Euphorbia milii Des Moul. var. tananarivae (Leandri) Ursch & Leandri

Euphorbia milii Des Moul. var. tulearensis Ursch & Leandri

Euphorbia milii Des Moul. var. vulcani (Leandri) Ursch & Leandri

Euphorbia milii Des Moul. var. milii

Euphorbia milii Des Moul. var. roseana Marn. -Lap. ex Demoly

Euphorbia milii Des Moul. var. splendens (Bojer ex Hook.) Ursch & Leandri

Euphorbia milii Des Moul. var. tenuispina Rauh & Razaf.

Euphorbia milii Des Moul. var. hislopii (N.E.Br.) Ursch & Leandri

Euphorbia millotii Ursch & Leandri

Euphorbia moratii Rauh var. moratii

Euphorbia moratii Rauh var. antsingiensis Cremers

Euphorbia moratii Rauh var. bemarahaensis Cremers

Euphorbia moratii Rauh var. multiflora Rauh

Euphorbia neobosseri (Rauh) Rauh var. itampolensis Rauh

Euphorbia neobosseri (Rauh) Rauh var. neobosseri

Euphorbia neohumbertii Boiteau var. aureoviridiflora Rauh = Euphorbia aureoviridiflora (Rauh) Rauh

Euphorbia neohumbertii Boiteau var. neohumbertii

Euphorbia pachypodioides Boiteau

Euphorbia parvicyathophora Rauh

Euphorbia paulianii Ursch & Leandri

Euphorbia perrieri Drake var. elongata Denis

Euphorbia perrieri Drake var. perrieri

Euphorbia platyacantha Drake = Euphorbia milii Des Moul. var. splendens (Bojer ex Hook.) Ursch & Leandri

Euphorbia primulifolia Baker var. begardii Cremers

Euphorbia primulifolia Baker var. primulifolia

Euphorbia primulifolia Baker var. isaloensis Rauh

Euphorbia quartziticola Leandri

Euphorbia razafindratsirae Lavranos

Euphorbia razafinjohanyi Ursch & Leandri

Euphorbia retrospina Rauh & Gerold

Euphorbia robivelonae Rauh

Euphorbia rossii Rauh & Buchloh

Euphorbia rubrostriata Drake var. doulioti Poiss.

Euphorbia rubrostriata Drake var. rubrostriata

Euphorbia sakarahaensis Rauh

Euphorbia splendens Bojer ex Hook. = Euphorbia milii Des Moul. var. splendens (Bojer ex Hook.) Ursch & Leandri

Euphorbia splendens Bojer ex Hook. var. betsileana Leandri

Euphorbia splendens Bojer ex Hook. var. bevilaniensis (Croizat) Leandri = Euphorbia milii Des Moul. var. bevilaniensis (Croizat) Ursch & Leandri

Euphorbia splendens Bojer ex Hook. var. breonii Leandri

Euphorbia splendens Bojer ex Hook. var. hislopii (N.E.Br.) Leandri = Euphorbia milii Des Moul. var. hislopii (N.E.Br.) Ursch & Leandri

Euphorbia splendens Bojer ex Hook. var. imperatae Leandri

Euphorbia splendens Bojer ex Hook. var. tananarivae Leandri

Euphorbia splendens Bojer ex Hook. var. vulcani Leandri

Euphorbia subapoda Baill. = Euphorbia primulifolia Baker var. primulifolia

Euphorbia suzannae-marnierae Rauh & Pétignat

Euphorbia tardieuana Leandri

Euphorbia thuarsiana Baill.

Euphorbia tsimbazazae Leandri

Euphorbia tulearensis (Rauh) Rauh

Euphorbia viguieri Denis var. ankarafantsiensis Ursch & Leandri

Euphorbia viguieri Denis var. capuroniana Ursch & Leandri

Euphorbia viguieri Denis var. tsimbazazae Ursch & Leandri

Euphorbia viguieri Denis var. viguieri

Euphorbia viguieri Denis var. vilanandrensis Ursch & Leandri

Euphorbia waringiae Rauh & Gerold

Euphorbia xanthadenia Denis = Euphorbia mahafalensis Denis var. xanthadenia (Denis) Leandri

Euphorbia × ambohipotsiensis Ursch & Leandri

Euphorbia × andrefandrovana Ursch & Leandri

Euphorbia × ingezalahiana Ursch & Leandri

Euphorbia × lomi Rauh

Euphorbia × mitsimbinensis Ursch & Leandri

Euphorbia × soanieranensis Ursch & Leandri

Euphorbia × zanaharensis Ursch & Leandri

Pedilanthus lycioides Baker = Euphorbia pedilanthoides Denis

Sterigmanthe bojeri (Hook.) Klotzsch & Garcke = Eu-

phorbia milii Des Moul.

Sterigmanthe splendens (Bojer ex Hook.) Klotzsch & Garcke = Euphorbia milii Des Moul. var. splendens (Bojer ex Hook.) Ursch & Leandri