Eusphalerum puberulum, Kiesenwetter, 1848

Herman, Lee H., 2001, Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the End of the Second Millennium. I. Introduction, History, Biographical Sketches, and Omaliine Group, Bulletin of the American Museum of Natural History 2001 (265), pp. 1-650 : 1-650

puberulum Kiesenwetter, 1848 , see: pallens Heer, 1841 .

pulcherrimum Bernhauer, 1901b: 654 ( Anthobium ; Type locality: Südtirol).

— Bernhauer and Schubert, 1910: 42 ( Anthobium ; catalog).

— Zanetti, 1982: 107 ( Eusphalerum ; characters; Italy).

— Zanetti, 1987: 115 ( Eusphalerum ; subgenus Eusphalerum ; characters; Italy).

— Zanetti, 1995: 6 ( Eusphalerum ; Italy).

- DISTRIBUTION: Italy.

pumilio Rosenhauer, 1856 , see: torquatum Marsham, 1802 .

punctatum Casey, 1894: 429 ( Anthobium ; Type locality: California: Santa Cruz Co.).

— Bernhauer and Schubert, 1910: 42 ( Anthobium ; catalog).

— Moore and Legner, 1975: 191 ( Eusphalerum ; checklist).

- DISTRIBUTION: USA.

puncticeps Luze, 1910 , see: torquatum Marsham, 1802 .

puncticolle Gredler, 1863 , see: minutum Fabricius, 1792 .

punctulatum Mulsant and Rey, 1880 , see: marshami Fauvel, 1869 .

ragusae Bernhauer, 1923 , see: luteum Marsham, 1802 .

ranunculi Gravenhorst, 1802 , see: minutum Fabricius, 1792 .

rectanguloides Smetana, 1967e: 298 ( Anthobium ; subgenus of Eusphalerum ; Type locality: Türkei: Erciyas dag, 2700 m).

— Zanetti, 1993: 219, 255 ( Eusphalerum ; characters; catalog; Turkey).

- DISTRIBUTION: Turkey.

rectangulum Baudi, 1870: 404 ( Anthobium ; [Note: Baudi cited this name as “ rectangulum Fauvel , in litt” so the later description of rectangulum by Fauvel may be the same species. Fauvel, 1871a: 78 referred to Baudi’s rectangulum as a subsequent reference of rectangulum Fauvel ]; Type locality: Alpi Grajis maritimi).

— Herman, 2001: 13 ( Eusphalerum ; synonymy).

- DISTRIBUTION: Europe.

rectangulum Fauvel, 1871: 17 ( Anthobium ; [preoccupied]; Type locality: Vosges; Alpes françaises; Piémont et Apennins ).

— Letzner, 1868: 360 ( Anthobium ; [Note: Misidentification: Bernhauer and Schubert, 1910: 42 cited sorbi Letzner as a synonym of rectangulum ; Letzner did not describe the species as new]; characters).

— Fauvel, 1871a: 78 [= 1872: 52] ( Eusphalerum ; [Note: Fauvel cited Baudi’s 1869 use of rectangulum as a subsequent use of rectangulum Fauvel ]; characters; France; Italy; Germany).

— Mulsant and Rey, 1880: 338 ( Anthobium ; characters; France).

— Ganglbauer, 1895: 752 ( Anthobium ; subgenus Eusphalerum ; characters; Germany; France; Alps; Apennines).

— Reitter, 1909: 196 ( Anthobium ; subgenus Eusphalerum ; characters; Germany).

— Bernhauer and Schubert, 1910: 42 ( Anthobium ; catalog).

— Portevin, 1929: 454 ( Anthobium ; characters; France).

— Scheerpeltz, 1933: 1036 ( Anthobium ; catalog).

— Freude, 1957: 107 ( Anthobium ; Germany).

— Coiffait, 1959c: 247 ( Eusphalerum ; subgenus Eusphalerum ; characters; France).

— Raianu, 1963: 85 ( Anthobium ; Romania).

— Horion, 1963: 40 ( Anthobium ; subgenus Eusphalerum ; Holland; Belgium; France; Switzerland; Italy; Corsica; Sardinia; Sicily; Romania; Czechoslovakia; Germany; Austria).

— Lohse, 1964: 32 ( Eusphalerum ; subgenus Eusphalerum ; characters; central Europe).

— Smetana, 1964d: 52 ( Anthobium ; subgenus Eusphalerum ; Czechoslovakia).

— Klinger, 1980: 456 ( Eusphalerum ; subgenus Eusphalerum ; characters of abdominal defensive gland).

— Tóth, 1982: 41 ( Eusphalerum ; characters; Hungary).

— Lucht, 1987: 86 ( Eusphalerum ; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Belgium; Netherlands; Luxembourg).

— Zanetti, 1987: 146 ( Eusphalerum ; subgenus Eusphalerum ; characters; Italy).

— Boháč, 1993: 40 ( Eusphalerum ; subgenus Eusphalerum ; Czech Republic; Slovakia).

— Zanetti, 1995: 6 ( Eusphalerum ; Italy).

— Herman, 2001: 20 ( Eusphalerum ; synonymic homonym of rectangulum ).

rectangulum Fauvel, 1871 , see: rectangulum Baudi, 1870 .

reitteri Bernhauer, 1935 a: 39 ( Anthobium ; Type locality: Ostsibirien: Sotka Gora).

— Tikhomirova, 1973a: 137 ( Eusphalerum ; subgenus Eusphalerum ; checklist; northern Siberia; Far East).

— Zanetti, 1993a: 49 ( Eusphalerum ; notes on characters and type locality; Siberia).

- DISTRIBUTION: Russia.

retowskii Bernhauer, 1914e: 33 ( Anthobium ; Type locality: Suchum, Caucasus, Gumista).

— Bernhauer, 1923d: 178 ( Anthobium ; characters; Caucasus).

— Scheerpeltz, 1933: 1036 ( Anthobium ; catalog).

— Tikhomirova, 1973a: 137 ( Eusphalerum ; subgenus Eusphalerum ; checklist; Caucasus).

— Zanetti, 1993: 217, 225, 247 ( Eusphalerum ; characters; catalog; Caucasus).

- DISTRIBUTION: Caucasus.

rhododendri Motschulsky, 1857 , see: anale Erichson, 1840 .

rhododendri Baudi, 1848: 148 ( Anthobium ; Type locality: Alpi Pedemontio; [Note: See lectotype designation by Zanetti, 1986]).

— Kraatz, 1857d: 1018 ( Anthobium ; synonym of sorbi ).

— Fauvel, 1871a: 79 [= 1872: 53] ( Eusphalerum ; characters; France; Switzerland; Italy).

— Mulsant and Rey, 1880: 339 ( Anthobium ; characters; France).

— Ganglbauer, 1895: 752 ( Anthobium ; subgenus Eusphalerum ; characters; western Alps).

— Reitter, 1909: 196 ( Anthobium ; subgenus Eusphalerum ; characters; Germany).

— Bernhauer and Schubert, 1910: 42 ( Anthobium ; catalog).

— Portevin, 1929: 454 ( Anthobium ; characters; France).

— Scheerpeltz, 1933: 1037 ( Anthobium ; catalog).

— Freude, 1957: 112 ( Anthobium ; Germany).

— Coiffait, 1959c: 247 ( Eusphalerum ; subgenus Eusphalerum ; characters; Jura; Alps).

— Horion, 1963: 40 ( Anthobium ; subgenus Eusphalerum ; France; Italy; Switzerland; Austria; Bosnia; Germany).

— Lohse, 1964: 32, 33 ( Eusphalerum ; subgenus Eusphalerum ; characters; central Europe).

— Zanetti, 1980b: 623 ( Eusphalerum ; Italy).

— Tóth, 1982: 42 ( Eusphalerum ; characters).

— Zanetti, 1986: 96 ( Eusphalerum ; lectotype designation).

— Zanetti, 1987: 146 ( Eusphalerum ; subgenus Eusphalerum ; characters; Italy).

— Lucht, 1987: 86 ( Eusphalerum ; checklist; Germany; Austria; Switzerland).

— Haghebaert, Bruge, and Drugmand, 1990: 35 ( Eusphalerum ; Belgium).

— Zanetti, 1995: 6 ( Eusphalerum ; Italy).

- DISTRIBUTION: Belgium, France, Germany, Switzerland, Italy.

obliquum Mulsant and Rey, 1861: 184 [= 1862: 168] ( Anthobium ; Type locality: Suisse).

— Fauvel, 1865c: 18 [= 1866: 15] ( Anthobium ; synonym of rhododendri ).

— Fauvel, 1871a: 79 [= 1872: 53] ( Eusphalerum ; synonym of rhododendri ).

— Mulsant and Rey, 1880: 340 ( Anthobium ; characters; France).

— Ganglbauer, 1895: 752 ( Anthobium ; synonym of rhododendri ).

— Bernhauer and Schubert, 1910: 42 ( Anthobium ; synonym of rhododendri ).

— Zanetti, 1987: 146 ( Eusphalerum ; synonym of rhododendri ).

bonadonai Coiffait, 1959c: 247 ( Eusphalerum ; subgenus Eusphalerum ; Type locality: Alpes Maritimes: Haute Tinée: Le Pré).

— Zanetti, 1986: 94 ( Eusphalerum ; synonym of rhododendri ).

— Zanetti, 1987: 146 ( Eusphalerum ; synonym of rhododendri ). rizzottivlachi Zanetti, 1992: 350 ( Eusphalerum ; Type locality: Tukalska Bjelina, Erzegovina meridionale).

- DISTRIBUTION: Bosnia Hercegovina.

robustum Heer, 1839: 179 ( Omalium ; Type locality: Basel).

— Fairmaire and Laboulbène, 1856: 646 ( Omalium ; characters; France).

— Kraatz, 1857d: 1010 ( Anthobium ; synonym of excavatum ).

— Fauvel, 1871a: 69 [= 1872: 43] ( Eusphalerum ; characters; France; Germany; Switzerland).

— Mulsant and Rey, 1880: 294 ( Anthobium ; characters; France).

— Ganglbauer, 1895: 749 ( Anthobium ; subgenus Eusphalerum ; characters; Alps; Pyrenees).

— Reitter, 1909: 196 ( Anthobium ; subgenus Eusphalerum ; characters; Germany).

— Bernhauer and Schubert, 1910: 42 ( Anthobium ; catalog).

— Portevin, 1929: 452 ( Anthobium ; characters; France).

— Scheerpeltz, 1933: 1037 ( Anthobium ; catalog).

— Freude, 1957: 112 ( Anthobium ; Germany).

— Coiffait, 1959c: 231 ( Eusphalerum ; subgenus Eusphalerum ; characters; Alps; Pyrenees; Massif Central).

— Horion, 1963: 49 ( Anthobium ; subgenus Eusphalerum ; France; Switzerland; Italy; Bosnia; Germany; Austria).

— Lohse, 1964: 34 ( Eusphalerum ; subgenus Eusphalerum ; characters; central Europe).

— Klinger, 1980: 456 ( Eusphalerum ; subgenus Onibathum ; characters of abdominal defensive gland).

— Zanetti, 1987: 120 ( Eusphalerum ; subgenus Eusphalerum ; characters; Italy).

— Lucht, 1987: 86 ( Eusphalerum ; checklist; Germany; Austria; Switzerland; France).

— Zanetti, 1995: 6 ( Eusphalerum ; Italy).

— Assing, Frisch, Kahlen, et al., 1998: 122 ( Eusphalerum ; character; Alps; Black Forest).

- DISTRIBUTION: France, Germany, Switzerland, Austria, Italy, Bosnia Herzegovina. alpestre Heer, 1839: 178 ( Omalium ; Type locality: Berglialp; Mühlebachalp).

— Fauvel, 1871a: 69 [= 1872: 43] ( Eusphalerum ; synonym of robustum ).

— Mulsant and Rey, 1880: 295 ( Anthobium ; synonym of robustum ).

— Ganglbauer, 1895: 749 ( Anthobium ; synonym of robustum ).

— Bernhauer and Schubert, 1910: 43 ( Anthobium ; synonym of robustum ).

excavatum Erichson, 1840: 893 ( Anthobium ; Type locality: Helvetia).

— Heer, 1841: 568 ( Omalium ; synonym of robustum ).

— Redtenbacher, 1849: 754 ( Anthobium ; characters; Austria).

— Redtenbacher, 1857: 255 ( Anthobium ; characters; Austria).

— Kraatz, 1857d: 1009 ( Anthobium ; characters; Germany).

— Fauvel, 1871a: 69 [= 1872: 43] ( Eusphalerum ; synonym of robustum ).

— Redtenbacher, 1874: 280 ( Anthobium ; characters; Austria).

— Mulsant and Rey, 1880: 295 ( Anthobium ; synonym of robustum ).

— Ganglbauer, 1895: 749 ( Anthobium ; synonym of robustum ).

— Bernhauer and Schubert, 1910: 43 ( Anthobium ; synonym of robustum ).

— Scheerpeltz, 1933: 1037 ( Anthobium ; synonym of robustum ).

— Zanetti, 1987: 120 ( Eusphalerum ; synonym of robustum ).

romani Coiffait, 1964 , see: limbatum Erichson, 1840 .

ruffoi Scheerpeltz, 1956: 3 ( Anthobium ; subgenus Eusphalerum ; Type locality: Gebiet des Mte. Pollino, teils Piani Pollino).

— Zanetti, 1982: 134 ( Eusphalerum ; characters; Italy).

— Zanetti, 1987: 131 ( Eusphalerum ; subgenus Eusphalerum ; characters; Italy).

— Zanetti, 1995: 6 ( Eusphalerum ; Italy).

- DISTRIBUTION: Italy.

rufipenne Gerhardt, 1910 [Note: Nomen nudum], see: primulae Stephens, 1834 .

rufiventre Cameron, 1930 , see: solitare Sharp, 1874 .

rufoscutellatum Eppelsheim, 1881: 508 ( Anthobium ; Type locality: Suramgebirge).

— Bernhauer and Schubert, 1910: 43 ( Anthobium ; catalog).

— Tikhomirova, 1973a: 137 ( Eusphalerum ; subgenus Eusphalerum ; checklist; Caucasus).

— Zanetti, 1993: 221, 250 ( Eusphalerum ; characters; catalog; Caucasus).

- DISTRIBUTION: Georgia.

gracile Luze, 1910: 241 ( Anthobium ; Type locality: Kaukasus, Suram).

— Scheerpeltz, 1933: 1034 ( Anthobium ; catalog).

— Zanetti, 1993: 250 ( Eusphalerum ; synonym of rufoscutellatum ).

rufotestaceum Motschulsky, 1860: 543 ( Anthobium ; Type locality: Alpes du Caucase; [Note: See lectotype designation by Zanetti, 1993]).

— Hochhuth, 1862: 111 ( Anthobium ; characters; Caucasus).

— Bernhauer and Schubert, 1910: 43 ( Anthobium ; catalog).

— Luze, 1910: 228 ( Anthobium ; notes).

— Scheerpeltz, 1933: 1037 ( Anthobium ; catalog).

— Tikhomirova, 1973a: 137 ( Eusphalerum ; subgenus Eusphalerum ; checklist; Turkey; Caucasus). — Zanetti, 1993: 217, 242 ( Eusphalerum ; characters; lectotype designation; catalog; Caucasus).

- DISTRIBUTION: Turkey, Caucasus.

rufulum Luze, 1910 [Note: Nomen nudum], see: viertli Ganglbauer, 1895 .

rugulosum Mäklin, 1853: 199 ( Anthobium ; Type locality: insula Sitkha).

— Fauvel, 1878d: 199 [= 1878e: 35] ( Anthobium ; characters; Sitka Island).

— Bernhauer and Schubert, 1910: 43 ( Anthobium ; catalog).

— Moore and Legner, 1975: 191 ( Eusphalerum ; checklist).

- DISTRIBUTION: USA.

ryvkini Zanetti, 1993: 221 , 225, 230, 253 ( Eusphalerum ; Type locality: near Teberda, Hatipara Mt., 1800 m).

- DISTRIBUTION: Russia.

sagiri Watanabe, 1990: 203 ( Eusphalerum ; subgenus Eusphalerum ; Type locality: Japan: Honshu: Gunma Pref.: near Kirizumi).

- DISTRIBUTION: Japan.

sahlbergi Luze, 1910: 244 ( Anthobium ; Type locality: Libanon, Monte Baruk ).

— Scheerpeltz, 1933: 1037 ( Anthobium ; catalog).

— Zanetti, 1993: 224, 261 ( Eusphalerum ; cited as shalbergi; characters; catalog; Lebanon).

- DISTRIBUTION: Lebanon.

saigokuense Watanabe, 1990: 191 ( Eusphalerum ; subgenus Eusphalerum ; Type locality: Japan: Honshu: Hiroshima Pref.: Haigamine).

- DISTRIBUTION: Japan.

sanae Watanabe, 1990: 211 ( Eusphalerum ; subgenus Eusphalerum ; Type locality: Japan: Honshu: Shizuoka Pref.: near Abe-tôge).

- DISTRIBUTION: Japan.

sareptanum Eppelsheim, 1878 a: 423 ( Anthobium ; subgenus Eusphalerum ; Type locality: Sarepta; [Note: See lectotype designation by Zanetti, 1993]).

— Bernhauer and Schubert, 1910: 43 ( Anthobium ; catalog).

— Scheerpeltz, 1933: 1037 ( Anthobium ; aberration of rufotestaceum ).

— Zanetti, 1993: 216, 222, 245 ( Eusphalerum ; lectotype designation; valid species; characters; catalog; Caucasus , Armenia, Turkey) .

- DISTRIBUTION: Armenia, Russia, Turkey.

starcki Reitter, 1890: 190 ( Anthobium ; Type locality: Cirkassien: Chag).

— Bernhauer and Schubert, 1910: 43 ( Anthobium ; synonym of sareptanum ).

— Scheerpeltz, 1933: 1037 ( Anthobium ; synonym of rufotestaceum ).

— Zanetti, 1993: 245 ( Eusphalerum ; synonym of sareptanum ).

umbricolor Roubal, 1915 a: 62 ( Anthobium ; Type locality: Caucasus occidentalis: Krasnaja Poljana; [Note: See lectotype designation by Zanetti, 1993]).

— Scheerpeltz, 1933: 1038 ( Anthobium ; catalog).

— Zanetti, 1993: 245 ( Eusphalerum ; lectotype designation; synonym of sareptanum ).

satanas Luze, 1911 [Note: Nomen nudum], see: tempestivum Erichson, 1840 .

satsuki Watanabe, 1990: 201 ( Eusphalerum ; subgenus Eusphalerum ; Type locality: Japan: Honshu: Yamanashi Pref.: Aokigahara, foot of Mt. Fuji).

- DISTRIBUTION: Japan.

schatzmayri C. Koch, 1938: 138 ( Anthobium ; Type locality: Monte Pollino, Appennino calabrese; [Note: See lectotype designation by Zanetti, 1980]).

— Zanetti, 1980: 167 ( Eusphalerum ; lectotype designation: M. Pollino, 1800 m; characters; Italy). — Zanetti, 1987: 113 ( Eusphalerum ; subgenus Eusphalerum ; characters; Italy).

— Zanetti, 1995: 6 ( Eusphalerum ; Italy).

- DISTRIBUTION: Italy.

alutaceum Scheerpeltz, 1956: 1 ( Anthobium ; subgenus Anthobium ; Type locality: Massiv des Mte. Pollino, Colle Gaudolino ).

— Zanetti, 1980: 167 ( Eusphalerum ; synonym of schatzmayri ).

— Zanetti, 1987: 113 ( Eusphalerum ; synonym of schatzmayri ).

schoenmanni Zanetti, 1993 a: 55 ( Eusphalerum ; Type locality: Seitengraben des Parwja Rietschka Tales, nördl. Wladiwostok).

- DISTRIBUTION: Russia.

scribae Schaufuss, 1862: 147 ( Anthobium ; Type locality: Südspanien).

— Bernhauer and Schubert, 1910: 43 ( Anthobium ; catalog).

— C. Koch, 1940: 373 ( Anthobium ; Spain).

— Coiffait, 1959c: 234 ( Eusphalerum ; subgenus Eusphalerum ; characters; Spain).

- DISTRIBUTION: Spain.

obscurum Brisout, 1866: 361 ( Anthobium ; Type locality: Espagne: environs de Madrid).

— Fauvel, 1869: 494 ( Anthobium ; synonym of scribae ).

— Bernhauer and Schubert, 1910: 43 ( Anthobium ; synonym of scribae ).

scutellare Erichson, 1840 , see: torquatum Marsham, 1802 .

segmentarium Mäklin, 1852: 322 ( Omalium ; Type locality: insula Sitkha).

— Fauvel, 1878d: 200 [= 1878e: 36] ( Anthobium ; characters; Alasaka).

— Bernhauer and Schubert, 1910: 43 ( Anthobium ; catalog).

— Moore and Legner, 1975: 191 ( Eusphalerum ; checklist).

- DISTRIBUTION: USA.

semicoleoptratum Panzer, 1795: 103 ( Dermestes ; Type locality: Not cited, presumably Germany).

— Panzer, 1799c: 6 ( Dermestes ; characters; illustrations).

— Gravenhorst, 1806: 219 ( Omalium ; synonym of testaceum ).

— Olivier, 1811: 480 ( Omalium ; characters; France).

— Lacordaire, 1835: 489 ( Omalium ; synonym of testaceum ).

— Stephens, 1839: 425 ( Anthobium ; synonym of torquatum ).

— Erichson, 1840: 893 ( Anthobium ; characters; Germany).

— Redtenbacher, 1849: 754 ( Anthobium ; synonym of abdominale ).

— Kraatz, 1857d: 1007 ( Anthobium ; synonym of abdominale ).

— Mulsant and Rey, 1880: 322 ( Anthobium ; synonym of abdominale ).

— Ganglbauer, 1895: 750 ( Anthobium ; synonym of abdominale ).

— Bernhauer and Schubert, 1910: 38 ( Anthobium ; synonym of abdominale ).

— Burakowski, Mroczkowski, and Stefańska, 1979: 24 ( Eusphalerum ; cited as valid name; catalog; Poland).

— Boháč, 1993: 40 ( Eusphalerum ; subgenus Eusphalerum ; Czech Republic; Slovakia).

— Ádám, 1996: 237 ( Eusphalerum ; valid name; Hungary).