Facitrichophora panama, Mathis, Wayne N. & Zatwarnicki, Tadeusz, 2012

Mathis, Wayne N. & Zatwarnicki, Tadeusz, 2012, A revision of the new world species of Polytrichophora Cresson and Facitrichophora, new genus (Diptera, Ephydridae), ZooKeys 231, pp. 1-116 : 12-14

Facitrichophora panama sp. n. Figs 1923

Diagnosis.

This species is distinguished from congeners by the following combination of characters: Small to medium-sized small shore flies, body length 1.84-3.20 mm.

Head: Frons gray to brown, mesofrons gray, parafrons grayish brown. Antenna mostly grayish brown to yellow; basal flagellomere blackish on lateral surface and somewhat dorsally; arista bearing 5-6 dorsal rays. Face silvery to whitish gray, broadly and somewhat protruding, lacking bare strip adjacent to carina, bearing 2 rows of setulae, medial row with longer setulae, 5-6, lateral row of setulae shorter, immediately adjacent to parafacial; parafacial becoming distinctly wider posteroventrally, making gena high (about 0.25 eye height), bearing setulae; facial and parafacial microtomentum silvery white to golden yellowish white, sometimes not concolorous; gena relatively high, equal or greater than 1/4 eye height; eye-to-cheek ratio 0.25-0.44.

Thorax: Mesonotum subshiny, especially medially, generally grayish peripherally to shiny copperish to blue, with sparse, whitish to grayish microtomentum; a single pair of posterior dorsocentral setae; prescutellar acrostichal setae small but evident, inserted close together; scutellum obtuse triangular, posterior margin shallowly rounded in do rsal view, lateral margins essentially straight, bearing 2 primary scutellar along lateral margins but otherwise lacking larger setae between primary setae. Wing hyaline or uniformly faintly infuscate; costal vein ratio 0.54-0.75; M vein 0.58-0.68. Coxae, femora, and tibiae grayish to blackish gray; hindtibia yellowish at apices; tarsi yellowish.

Abdomen: Tergite 5 of male narrowed apically. Male terminalia (Figs 19-22): Epandrium connected dorsally above cerci (Figs 19-20), in posterior view polygonal oval, slightly wider at midheight, dorsal margin broadly rounded, ventral margin deeply incised, each side broadly and deeply bifurcate, lateral lobe narrowly triangular, apex narrowly rounded, more setulose toward dorsal and ventral ends, in lateral view with ventral apex bifurcate and bearing dense patch of setulae; cerci moderately large, conspicuous; gonite in lateral view rod-like, shallowly sinuous (Fig. 22), in ventral view (Fig. 21) semihemispherical; aedeagus in lateral view simple, elongate, tubular, irregularly tapered to narrow, digitiform process, apex truncate, in ventral view (Fig. 21) elongate, narrow, gradually tapered to ventral apex; phallapodeme with portion toward aedeagal base rod-like, very narrow and elongate, tapered with apical half parallel sided, hypandrial end bearing irregularly shaped, pointed keel, in ventral view (Fig. 21) an elongate rod, expanded at each end, end toward aedeagal base shallowly incised; gonite in lateral view (Fig. 22) elongate, very narrow, shallowly and angularly curved, in ventral view (Fig. 21) broadly zigzagged, narrower toward aedeagal base, becoming generally broader toward hypandrium; hypandrium in lateral view (Fig. 22) elongate, very narrow, shallowly curved, in ventral view (Fig. 21) more or less deeply V-shaped, base broad with rounded anterior margin, each posterior arm oriented posterolaterally, elongate, parallel-sided, pointed apically.

Type material.

The holotype male is labeled "PANAMA[.] DarienProv[incia][,] Garachine/Feb 1953[,] FSBlanton/USNM ENT 00285973 [plastic bar code label]/HOLOTYPE ♂ Facitrichophora panama Mathis & Zatwarnicki, USNM [red]." The holotype is double mounted (glued to a paper triangle), is in moderate condition (arista missing; some head setae missing or misoriented), and is deposited in the USNM. One male paratype (dissected; USNM) bears the same label data as the holotype. Other paratypes are as follows: PANAMA. Canal Zone: San Lorenzo (9°07.9'N, 79°32.1'W), 15 Aug 1952, F. S. Blanton (1♀; USNM). Darien: El Real (8°07.8'N, 77°43.6'W), 8 Aug 1952, F. S. Blanton (1♀; USNM). Los Santos: Bayano (7°57'N, 80°20'W), 8 Apr 1952, F. S. Blanton (1♀; USNM).

Type locality.

Panama. Darien: Garachine (8°04'N, 78°22'W).

Distribution

(Fig. 23). Neotropical: Panama (Canal Zone, Darien, Los Santos).

Etymology.

The species epithet, panama, refers to the country where this species was collected and is a noun in apposition.

Remarks.

Although similar to Facitrichophora manza (similarities include having a single dorsocentral seta, the mesonotum grayish brown to bronzish brown, and having two lateral scutellar setae), it is distinguished by having male tergites three and four subequal in size (tergite four not unusually longer as in Facitrichophora manza ); male tergite three is not produced ventrolaterally; and male tergite five is rounded or angulate (not projected anteroventrally and acutely pointed as in Facitrichophora manza ).