Festuca laevis subsp. marginata (Hack.) Fournier, 1928

López, Eusebio, Martínez-Sagarra, Gloria & Devesa, Juan Antonio, 2017, Taxonomic treatment of the Festuca marginata group (Pooideae, Poaceae) on the Iberian Peninsula (SW Europe), Phytotaxa 319 (3), pp. 201-224 : 217-221

- Panicle 2.2–9.5(14) cm, cylindrical, with 1–4(7) branches or not branched.............................................. b. subsp. andres-molinae a. subsp. marginata

Festuca ovina var. timbalii Hackel (1882: 108) . Festuca ovina subsp. timbalii (Hack.) Richter (1890: 96) . Festuca timbalii (Hack.) Kerguélen (1983: 61) . Festuca ovina var. timbalii (Hack.) Rouy (1913: 212) . Lectotype (designated by Kerguélen 1983):— FRANCE. Haute-Garonne: Caraman, August 1874, Timbal-Lagrave (W).

Festuca ovina var. gallica Hackel ex Charrel (1920: 5) . Festuca ovina subvar. hervieri St. -Yves in Litardière (1923: 105, 106). Festuca laevis subsp. gallica (Hackel ex Charrel) Fournier (1928: 587) . Festuca duriuscula var. gallica (Hackel ex Charrel) Fournier (1935: 80) . Festuca hervieri (St.-Yves) Patzke (1962: 213). Festuca marginata subsp. gallica (Hack. ex Charrel) Breistroffer (1963: 79) . Lectotype (designated by Kerguélen 1975):— FRANCE. Drôme: Romans, banks of Isère, calcareous hills, June 1894, Hervier (G!).

Lowest node 2–6 cm above base of plant. Tiller leaf blade 6–30 cm × 0.6–1.1 mm, auricles absent or 0.1–0.3 mm, with 7–9 vascular bundles. Uppermost cauline leaf 4.5–9 cm. Panicle 5–12 cm, condensed, ovate-oblong, with 4–9 branches. Spikelets 5.5–11 mm, green, yellowish, or slightly greenish-purple. Lemma of the basal flower muticous or with awn of 0.1–0.9 mm, that of the second basal flower muticous or with awn of 0.1–1.4 mm, and up to 1.8 mm in the rest.

Ecology and distribution:—Forest fringes and clearings, roadsides, and embankments, on calcareous substrates; 1250–1650 m; flowering from May to July. Pyrenees and most of France ( Fig. 4 ).

Additional specimens examined: — ANDORRA. Andorra La Vella, Arinsal , 5 July 1992, Montserrat & Benito s.n. (BC 878917, JACA 182492 ) . FRANCE. Ardèche: Near Mayres, Rocher d’Abraham , 1 July 1971, Auquier F 1114 (MA 286731, MAF 96011 ). Drôme: Romans, banks of Isère, June 1894, Hervier s.n. (G 164992) ; ídem, June 1897, Hervier s.n. ( MA 12082 ) ; Romans-sur-Isère, Les Balmes, 28 May 1972, Auquier et al. F1138 (MA 286731, MAF 98943 ). Meurthe-et-Moselle: Francheville, N of Toul, 5 May 1973, Auquier F 1189 (MA 363067, MAF 98942 ). Puy-de-Dôme: Chanturges, near Clermont-Ferrand, without date, Héribaud s.n. ( LE 9826 ). Seine-et-Marne: Chateau- Landon, Pont Franc, 2 June 1929, Chouard s.n. ( MA 466216 ). Yvelines: Gommécourt, 9 June 1980, De Retz 80379 (MA 378091, MAF 122776 ) . SPAIN. Gerona: Llivia , castle, 24 May 2014, E. López & G. Martínez GM110/14 ( COFC 62021 ) .

b. subsp. andres-molinae Fuente & Ortúñez (1993: 107). Holotype:— SPAIN. Burgos : Covarrubias, 23 June 1978, Izco & Molina (MAF 136501!)

Lowest node 1–4 cm above base of plant. Tiller leaf blade 1.5–13 cm × (0.3) 0.4–0.8 mm, auricles 0.1–0.4(0.6) mm, with 7(9) vascular bundles. Uppermost cauline leaf 1–4(5.5) cm. Panicle 2.2–9.5(14) cm, lax or condensed, cylindrical, with 1–4(7) branches or not branched. Spikelets 5–8.5(9.4) mm, green, yellowish, or slightly greenish-purple. Lemma of the basal flower muticous or with awn of 0.1–0.6 mm, that of the second basal flower muticous or with awn of 0.1–0.6 mm, and up to 1 mm in the rest.

Ecology and distribution:—Holm oak and Portuguese oak woods, scrub mesoxerophilic brome and grass lands on calcareous substrates; (180) 520–1550 m; flowering from May to August. Endemic to the NE and E Iberian Peninsula ( Fig. 4 ).

Additional specimens examined: — SPAIN. Álava: Pipaón, Pass of Recelle , July 1933, Losa s.n. ( MA 11934 ). Albacete: Bonete, Bonete - Alpera, Sierra del Mugrón , 29 May 2003, Crespo et al. s.n. ( ABH 47678 ). Barcelona: Alt Berguedà, La Clusa, crest of Baga de Sant Romà , 10 July 1976, Rosell s.n. ( BC 624959 ) ; Castanyadell , June 1869, Masferrer s.n. ( BC 69819 ) ; Montcau , 16 June 1946, De Bolòs s.n. ( BC 102112 ) ; Montserrat, top of Montgros , 2 May 2003, Pyke & Pallàs s.n. ( BC 905794 ) ; Serra de Pedraforca , crags of Gresolet, 1 August 1926, Font Quer & Cuatrecasas s.n. ( BC 868896 ). Burgos: Bujedo , 8 June 1924, Elías s.n. ( MA 11957 ) ; Covarrubias , 23 June 1978, Izco & Molina s.n. ( MAF 136501 ) ; Encio , 30 May 1906, Sennen & Elías s.n. (BC 75589, BC 86470, MA 59444) ; Hoyos del Tozo , 15 June 1985, Alejandre s.n. ( MA 339373 ) ; Los Altos, Tudanca, Cidad de Ebro , 3 June 2006, Benito Aparicio s.n. ( SALA 145422 ) ; Medina de Pomar - Santurde , 20 June 1992, Elorza et al. s.n. ( COFC 62160 ) ; Merindad de Cuesta - Urria, Mijangos, river Nela , 3 July 1998, Fernández & Patino s.n. ( COFC 62159 ) ; ídem, Quintana de Entrepeñas , 6 June 1993, Elorza & Patino s.n. ( COFC 62158 ) ; Montes Obarenes , 16 June 1912, Elías s.n. ( MA 11976 ) ; Pancorbo , 24 May 1909, Elías s.n. ( MA 59514 ) ; ídem, Barranco de Barbalante , 9 June 1914, Font Quer s.n. (BC 68895, MA 12088) ; Valle de las Caderechas, June , Losa s.n. (MA 12112, MA 466279). Gerona : Sant Feliu de Buixalleu , Castell de Montseriu , 30 May 1986, Vilar s.n. ( HGI 10065 ). La Rioja: around Logroño, without date, Zubia s.n. ( MA 11933 ) ; Nieva de Cameros , July 1905, Pau s.n. ( MA 59454 ) ; San Vicente de la Sonsierra, Peñacerrada - Rivas de Tereso, 16 July 1996, De la Fuente & Otúñez s.n. ( MAF 159650 ). Lérida : Montsec d’Ares , Colobó , 12 June 1980, Romo s.n. ( BC 915983 ) ; Vilanova de la Sal, 12 May 1933, Font Quer s.n. ( BC 111181 ). Navarra: Izánoz - San Miguel de Izaga, 12 June 2008, Sánchez & Ibáñez s.n. (ABH 53373, HUAL 19653, MA 806370, MAF 168302, SALA 136681 ). Palencia: Quintana del Puente , 13 June 1987, C. López & Romero Abelló s.n. ( MA 619548 ) ; ídem, 18 June 1988, C. López et al. s.n. ( MA 627237 ) ; ídem 15 July 1988, Carrasco et al. s.n. ( MA 627252 ) ; San Cebrián , 15 July 1989, Carrasco et al. s.n. ( MA 617529 ) ; Villamediana, near road to Astudillo, 15 July 1989, Carrasco et al. s.n. ( MA 617529 ). Segovia: Sebulcor, 12 June 1983, Romero s.n. ( SALA 40424 ). Soria: Silvestre de Navaleno , 14 September 1932, Villar s.n. ( MA 157790 ). Tarragona: Alfara de Carles , NW of Vall Cervera, 17 May 1986, De Torres s.n. ( BC 905474 ) ; Coll del Caragol , 8 June 1999, Navarro et al. CN–2373 ( MA 626894 ) ; Massis del Port, Tortosa, Barranc d’Aigua Donzella , 20 July 1987, De Torres s.n. ( BC 905497 ) ; ídem, Font del Caragol - Els Bassis, Tortosa , 26 June 1983, De Torres s.n. ( BC 905473 ) ; ídem, Les Clotes, Horta de Sant Joan , 16 June 1987, De Torres s.n. ( BC 905499 ) ; ídem, Millers , La Sénia, 26 June 1983, De Torres s.n. ( BC 905496 ) ; ídem, Tres Pins, Roquetes , June 1982, De Torres s.n. ( BC 905498 ) ; Monte Caro , without date, Font Quer s.n. ( BC 69748 ) ; Ports de Tortosa, Els Bassis, 18 June 1935, Font Quer & Rothmaler s.n. ( BC 636267 ). Teruel: Monreal del Campo , without date, Benedicto s.n. ( BC 69701 ). Valladolid: Encinas de Esgueva , Fuenteblanca, Cañada de Vadetorres , 14 June 1981, Fernández JF 472 ( MA 548287 ). Zaragoza: Dam of Yesa , Channel of Berdún , 30 June 1991, De la Fuente & Ortúñez s.n. ( MAF 159651 ) ; Calatayud , without date, Pyke s.n. ( BC 905816 ) ; ídem, Embid de la Ribera, Peña de la Mora, 29 May 2006, Pyke SBP5152 ( BC 905757 ) ; ídem, Sierra de Vicort , without date, Pyke s.n. ( BC 905817 ) ; Campillo de Aragón, Ribera de la Hoz, 25 June 2003, Ladero et al. s.n. ( SALA 115118 ) .

2. Festuca lambinonii Kerguélen (1983: 59) ( Fig. 5 ). Festuca ovina subvar. alopecuroides Hackel (1882: 108) . Festuca marginata subsp. alopecuroides (Hack.) Richter (1890: 96) . Festuca ovina var. alopecuroides (Hack.) Rouy (1913: 212) . Lectotype (designated by Kerguélen 1983: 59):— FRANCE. Pyrénées-Orientales: Prades, font de Comps, 7 July 1872, Timbal-Lagrave (W)

Festuca ovina subvar. platyphylla Saint-Yves (1924: 41) . Lectotype: not designated, probably in Saint-Yves’ Herbarium of Festuca housed in G ( Kerguélen 1983); Isolectotype in MPU ( Kerguélen 1983: 59):— FRANCE. Pyrénées-Orientales: Vallon de Ria, May 1897, Sennen.

Festuca rivas-martinezii Fuente & Ortúñez (1994: 25) . Festuca longifolia subsp. rivas-martinezii (Fuente & Ortúñez) Cebolla, López Rodr. & Rivas Ponce in Cebolla et al. (1997: 33). Holotype:— SPAIN. Madrid: Sierra de Guadarrama, pass of Navafría, 15 June 1989, Ortúñez & De la Fuente (MAF 137908!).

Festuca rivas-martinezii subsp. rectifolia Fuente, Ortúñez & Ferrero Lom. in Fuente et al. (1997: 319). Festuca rectifolia (Fuente, Ortúñez & Ferrero Lom.) Cebolla & Rivas Ponce (2003: 210) . Holotype:— SPAIN. Vitoria: Pipaón, pass of Herrera, Sierra de Cantabria, 16 July 1996, De la Fuente, Ortúñez & Ferrero 234 (MAF 152600!).

Festuca longifolia auct. non Festuca longifolia Thuillier (1799: 50) .

Festuca arvernensis subsp. costei auct. non Festuca costei (St.-Yves) Markgraf-Dannenberg (1978: 327).

Festuca burgundiana auct. non Festuca burgundiana Auquier & Kerguélen (1978: 63) .

Festuca marginata subsp. gallica auct. non Festuca marginata subsp. gallica (Hackel ex Charrel 1920: 5) Breistroffer (1963: 79) .

Perennial herb, tufted, culm up to 76 cm. Lowest node 1–6 cm above base of plant. Tiller leaf sheath with margins free or fused about 1/2 their length, ligule 0.1–0.5 mm, auricles absent or of 0.1–0.7 mm. Blades 1.5–25 cm × 0.4–1.3 mm, narrowly linear, curved, or lightly curved distally in a spiral, somewhat stiff and weakly prickled, conduplicate, glabrous, somewhat scabrid at the apex, usually pruinose, with 5–9 ribs and sclerenchyma in three thin strands, the median strand decurrent, rarely continuous or interrupted. Panicle 2.5–11(15.5) cm, condensed, ovate-oblong, with 1–8 branches. Spikelets 5–10.5(13) mm, with peduncle 0.5–6 mm, with (2)3–8(12) fertile flowers and frequently 1 sterile and rudimentary apical flower. Glumes narrowly scarious on the margins, acuminate, glabrous or ciliolate at the top margin, the lower 1.5–4 mm and o the upper 2–5 mm. Lemma 3.6–6 mm, obovate-lanceolate, acuminate, with narrowly scarious margins in at least the upper third, sometimes ciliate, glabrous or scabrid distally on the back, that of the basal flower with a mucro or awn of 0.1–1.5(1.8) mm, that of the second basal flower with an awn of (0.3)0.4– 3(3.5) mm, and up to 3.5 mm in the rest. Anthers 1.7–2.9(3.1) mm. Ovary glabrous. Caryopsis 2.5–2.8 mm.

Ecology and distribution:—Montane and subalpine herb and grasslands, rocky areas, roadsides and cleared Pyrenean oak, pine, and Spanish juniper woodlands, and abandoned cropland, on siliceous or calcareous substrates; 800–2300 m; flowering from (April) May to September. C, N and E of the Iberian Peninsula ( Andorra, Spain and France) ( Fig. 4 ).

Additional specimens examined: — ANDORRA. Andorra La Vella, 20 May 1992, P. Montserrat s.n. ( BC 878916 ) ; Cortal d’Encamp , 10 July 1981, Van Buggenhout s.n. ( MAF 160324 ) ; Pass of Envalira, Pic Maia , 4 May 2014, E. López & G. Martínez GM117/14 ( COFC 61994 ) . FRANCE. Pyrénées-Atlantiques: Lescun, Cayolar d’Anaye , 12 July 1998, Patino & Valencia s.n. (COFC 62171, COFC 62751 ) . SPAIN. Álava: Peñacerrada, Puerto de Herrera - Peñacerrada , 16 July 1996, De la Fuente et al. s.n. ( MAF 159665 ) ; Pipaón, Pass of Herrera, 16 July 1996, De la Fuente et al. s.n. ( MAF 152600 ). Albacete: Higueruela, El Molatón, 28 May 2003, Crespo et al. s.n. ( ABH 47552 ). Alicante: Confrides, Sierra de la Serrella, Penyes del Albardar , 24 June 1993, Solanas s.n. ( ABH 19335 ) ; La Torre de les Maçanes, Sierra del Rentonar , 8 June 1994, Serra et al. s.n. ( ABH 10221 ) ; Parcent , Sierra del Carrascal, 11 June 1997, Signes & Soler 7203–JXS ( MA 590188 ) ; Quatretondeta, Hort dels Frares , 17 June 1989, Solanas s.n. (ABH 4146, MA 534054) ; Sierra de Aitana , 9 July 1958, Fabregat s.n. ( ABH 23333 ) ; ídem, Alcoleja, Font del Espinar , 23 July 1989, Solanas s.n. ( ABH 4149 ) ; ídem, Benifato, Font de Forata , 18 June 1992, Solanas s.n. (ABH 4148, ABH 4152 ) ; ídem, 6 July 1993, Solanas s.n. ( ABH 7905 ) ; ídem, Font de Partagat , 22 June 2005, Alonso et al. s.n. ( ABH 49511 ) ; ídem, Font de Partagat , Pas de la Rabosa, 25 May 2014, Alonso et al. GM01/14 ( COFC 61532 ) ; ídem, 2 July 1992, Solanas s.n. (ABH 4147, ABH 4150, MAF 141882 ) ; ídem, 9 July 1972, Fabregat s.n. ( ABH 23332 ) ; ídem, Penya Alta , 28 May 1992, Solanas et al. s.n. ( ABH 3990 ) ; Vall de Laguart, Sierra de la Carrasca , 14 June 1997, Signes & Soler 7303–JXS ( MA 590378 ) ; ídem, Sierra del Peñón, Benimaurell , 22 May 1993, Soler 579 ( MA 548676 ) ; Villena, Sierra de Salinas , 4 July 1993, Rigual et al. s.n. ( ABH 6484 ). Ávila: Garganta del Villar - Navacepedilla de Corneja , Pass of Chía , 4 July 2012, Devesa & G. Martínez JADA601–3 ( COFC 61504 ) ; Hoyocasero, Pinar de Hoyocasero , 15 June 2013, Luceño & Maguilla s.n. ( UPOS 5285 ) ; La Serrota, Canto de la Oración, Sierra de la Paramera, 22 June 1995, De la Fuente & Ferrero s.n. ( MAF 159659 ) ; Puerto del Pico , 23 June 2015, Devesa & G. Martínez JADA696B ( COFC 62035 ) ; Sierra de Ojos Albos, Los Regajales , 3 July 1984, Burgaz et al. s.n. (MA 408110, MA 411398, MA 411422) ; Valle de Amblés, Puerto de Menga , 26 July 1976, Fuertes Lasala s.n. ( MA 213638 ) ; Villanueva del Campillo, 21 May 1995, De la Fuente s.n. ( MAF 159660 ). Barcelona: Bagà - Coll de Pal , 5 July 2003, Pyke s.n. (BC 863692, BC 863694, BC 863695) ; Coll de Pal , 5 July 2003, Pyke s.n. (BC 863693, BC 863696, BC 863697) ; idem, between Bagà and the pass, 5 July 2003, Pyke s.n. ( BC 863691 ). Burgos: Contreras, Alto de San Carlos , 18 July 1979, Muñoz Garmendia et al. 507 ( MA 413100 ) ; Gamonal , 16 June 1914, Font Quer s.n. ( MA 59487 ) ; Miranda, Monte de Quintana , without date, Losa s.n. ( MA 59462 ) ; Miranda, without date, Losa s.n. ( MA 146418 ) ; Pineda de la Sierra, 26 June 1914, Font Quer s.n. ( MA 59486 ) ; ídem, 5 September 1998, Ferrero et al. s.n. ( MAF 159662 ) ; ídem, valley of Carmina Mine , 29 June 2006, Benito Aparicio s.n. ( SALA 145420 ) ; Quintanabaldo - Puentedey, 5 June 2004, Benito Alonso s.n. ( SALA 124554 ). Cantabria: Serna, Hermandad de Campóo de Suso , 6 July 1983, Aedo s.n. ( MA 615675 ). Castellón: Algimia de Almonacid , 20 June 2012, Alonso et al. s.n. ( ABH 69212 ) ; ídem, Sierra de Espadán , 19 June 2012, Alonso et al. s.n. ( ABH 69189 ) ; Penyagolosa , July 1962, Vigo s.n. ( BC 602321 ) ; Sacañet, Navas de Torrijas , 8 July 1895, Pau s.n. ( MA 12033 ) ; Sant Joan de Penyagolosa , El Rebollar, 13 July 1962, Vigo s.n. ( BC 602328 ) ; Vistabella del Maestrat , 25 June 1963, Vigo s.n. ( BC 602323 ) ; ídem, July 1963, Vigo s.n. (BC 602329, BC 602332, BC 602335) ; Vistabella del Maestrat - Tossal Marinet, July 1963, Vigo s.n. ( BC 602322 ). Cuenca: Beteta , 13 July 1932, Caballero s.n. ( MA 146746 ) ; Serrania de Cuenca, Uña , June 1962, Borja s.n. ( MAF 75939 ) ; Sierra de Valdemeca , 20 July 1974, G. López s.n. ( SALA 56625 ) ; Forest areas of Alto Júcar, 29 June 1966, Rivas Goday & Borja s.n. ( MAF 75095 ). Gerona : Alp , 7 June 1993, Soriano & Ninot s.n. ( MAF 143230 ) ; Arbúcies, Espinalves , 25 May 2003, Pyke s.n. ( BC 905799 ) ; Arbucíes - Sant Hilari , 21 May 2012, Devesa & E. López 582B ( COFC 61171 ) ; Coll d’Ares , 23 May 2014, E. López & G. Martínez GM95/14 (COFC 61581, COFC 61582 ) ; La Molina, Alp, Tosa d’Alp , 9 July 2006, Romo et al. s.n. ( BC 866330 ) ; Maçanet de la Selva, Empalme , 31 May 1945, Font Quer s.n. ( BC 868850 ) ; Olopte - Tossal d’Isovol , 28 June 2006, Romo & Nualart R-14130 ( BC 866340 ) ; Ribes de Freser , 14 February 2003, Pyke s.n. ( BC 905798 ) ; Vall de Nuria, Pico Noufont - Sanctuary , 23 July 2014, E. López & G. Martínez GM109/14 ( COFC 61578 ) ; Vall de Ribes, Angelats , 3 July 1971, Vigo s.n. ( BC 612913 ) ; ídem, 4 September 1972, Vigo s.n. ( BC 612929 ) ; ídem, Bosc de Ribes , 24 July 1972, Vigo s.n. ( BC 612921 ) ; ídem, Campelles , 14 August 1972, Vigo s.n. ( BC 612890 ) ; ídem, 3 July 1971, Vigo s.n. (BC 612918, BC 612922) ; ídem, Capdevila , 18 June 1970, Vigo s.n. ( BC 612928 ) ; ídem, Gorges del Freser , 21 July 1972, Vigo s.n. ( BC 612914 ) ; ídem, Les Guilleumes , 1 July 1972, Vigo s.n. ( BC 612912 ) ; ídem, Pedrisses , 25 July 1970, Vigo & Anglada s.n. ( BC 612925 ) ; ídem, Planoles , 21 June 1970, Vigo & Anglada s.n. ( BC 612931 ) ; ídem, Planoles - Dòrria , 10 July 1972, Vigo s.n. ( BC 612915 ) ; ídem, Queralbs , 17 June 1971, Vigo s.n. ( BC 612926 ) ; ídem, 24 June 1971, Vigo & Anglada s.n. (BC 612920, BC 612923) ; ídem, Queralbs, Torrent del Forn, 21 July 1972, Vigo s.n. ( BC 612916 ). Guadalajara: Cantalojas , Valle del Lillas , 11 June 1986, Burgos & Cardiel s.n. ( MA 498996 ) ; Checa, Fuente del Hocinillo, 21 May 1995, Martín Ballesteros 1120 ( SALA 59938 ). Huesca : Formigal , 4 July 2013, Devesa & G. Martínez GM72/13 ( COFC 61460 ) ; La Peña, 20 May 1998, Pyke s.n. ( BC 905815 ). León: Sena de Luna, La Collada de Aralla, 17 July 1995, De la Fuente & Ferrero s.n. ( MAF 159663 ) ; Torre de Babia, 29 June 1991, Gómiz s.n. ( MA 569900 ). Lérida: Bossost, Vall de Aran , 8 June 1874, Compañó s.n. ( BC 610263 ) ; Espot, Ribera de Estanyets , 10 July 1934, Rothmaler s.n. ( BC 78532 ) ; ídem, Ribera de Peguera , 12 July 1934, Rothmaler s.n. ( BC 78531 ) ; Monteixo, Àreu , 2 August 1911, Font Quer s.n. ( BC 69860 ) ; Serra del Cadí, Prat d’Aguiló , 9 July 1949, De Bolòs et al. s.n. ( BC 822761 ) ; Vall d’Aran, without date, Costa s.n. ( BC 616728 ). Madrid: Cadalso de los Vidrios , July 1956, Borja s.n. ( MA 201986 ) ; top of Las Guarramillas , August 1911, Beltrán s.n. ( MA 59467 ) ; Lozoya, Pass of Navafría , 20 June 1998, Ferrero s.n. ( MAF 159661 ) ; Miraflores de la Sierra, June 1961, Borja s.n. ( MA 201987 ) ; Monte de Valdelatas , without date, Cutanda s.n. (MA 11918, MA11919) ; Montejo de la Sierra, Hayedo de Montejo , 6 June 2012, Manzano s.n. (MAF 174093, MAF 174097, MAF 174098, MAF 174099, MAF 174100, MAF 174101 ) ; Pico Abantos , June 1988, Ortúñez s.n. ( MAF 159664 ) ; Pass of Canencia , 12 June 1999, Rojo s.n. ( MAF 160654 ) ; ídem, 24 June 2015, Devesa & G. Martínez JADA702B ( COFC 62039 ) ; ídem, 28 May 2013, G. Martínez & C. Morales s.n. ( COFC 62012 ) ; ídem, 9 July 1956, without collector s.n. ( MA 548773 ) ; ídem, El Rasero, 9 July 1956, without collector s.n. (MA 548787, MA 548790) ; Pass of Navafría, 15 June 1989, Ortúñez & De la Fuente s.n. ( MAF 137908 ) ; Rascafría, Pass of Morcuera , 14 June 2014, G. Martínez & C. Morales GM37/14 ( COFC 61476 ) ; Robregordo , 19 June 1918, C. Vicioso s.n. ( MA 11960 ) ; Sierra de Guadarrama, Alto de los Leones , 14 June 2014, E. López EL03/14 ( COFC 61498 ) ; ídem, June 1964, Borja s.n. (MA 195514, MAF 66348, MAF 72158, SEV 6310 ) ; Sierra de Guadarrama , June 1958, A. Rodríguez s.n. ( MA 186145 ) ; ídem, July 1963, Borja s.n. ( MA 201988 ) ; ídem, June 1964, Borja & A. Rodríguez s.n. ( MA 186122 ) ; Pass of Guadarrama, without date, Cutanda s.n. ( MA 11962 ). Segovia : Cerezo de Arriba , 31 May 2002, Rojo s.n. ( MAF 163831 ) ; ídem, Peak of El Lobo , 18 June 1983, T. Romero s.n. ( MA 569177 ) ; La Granja de San Ildenfonso - Torrecaballeros , 21 June 1985, R. García 989 ( MA 837393 ) ; Navares de las Cuevas, Peñacuerno , 27 June 1984, T. Romero s.n. ( MA 569262 ) ; Navas del Espinar , 13 July 1974, Ladero & Rivas Goday s.n. ( SALA 23147 ) ; Prádena , 7 June 1987, García Adá 3725RG ( MA 837394 ) ; Pass of La Quesera , Hayedo de la Pedrosa, 7 July 1985, Luceño et al. s.n. ( MA 373324 ) ; Pass of Los Leones , N face, 4 July 1975, Fernández Casas 507 ( MA 412010 ) ; San Rafael , 14 June 1924, Villar s.n. ( MA 157793 ) ; Sierra de Guadarrama, Valsaín , July 1916, Vicioso s.n. (MA 11951, MA 11952) ; Siguero , 4 June 1985, T. Romero s.n. ( MA 569179 ) ; Torreadrada, Fuente Merina , 27 June 1984, T. Romero s.n. ( MA 569263 ) ; Somosierra, without date, Cuatrecasas s.n. ( MA 11966 ). Soria: Montenegro de Cameros , June, Caballero s.n. ( MA 59383 ) ; ídem, Pico Berezoles , 20 July 2005, Molina & Montamarta s.n. ( MA 737470 ) ; Soria, eastern slopes of the Castle, 10 June 1946, Montserrat s.n. ( BC 112329 ). Teruel: Sierra de Gúdar , Valdelinares , August 1954, without collector s.n. ( MA 186119 ) ; Sierra de Javalambre, 9 July 1895, Pau s.n. ( BC 69771 ). Valencia: Cortes de Pallás , 5 June 2003, Ferrer & Guara 03/1294 (BC 81272, BC 82558). Zaragoza: Sierra del Moncayo , between parking and the Sanctuary of Moncayo , 30 July 2013, Devesa & G. Martínez GM49/13 (COFC 61499, COFC 61501, COFC 61502, COFC 61503, COFC 61505, COFC 61509 ) ; ídem, Sanctuary , 10 July 1994, Aedo et al. CA3122 ( MA 544194 ) ; Tarazona, Moncayo, Pass of Castilla , 19 June 2002, P. García & Linares s.n. ( BC 862861 ) .

Figures

FIGURE 4. Distribution map of Festuca marginata subsp. marginata, F.marginata subsp. andres-molinae and F.lambinonii in the Iberian Peninsula.

FIGURE 5. Festuca lambinonii (SPAIN. Gerona: Coll d’Ares, 23 May 2014, E. López & G. Martínez; COFC 61582):A. Habit. B. Detail of the apex of the sheath and ligule in adaxial view. C. Spikelet. D. Lower glume in abaxial view. E. Upper glume in abaxial view. F. Lemma in abaxial view. G. Palea in abaxial view. H. Lodicule. I. Sexual whorls. J. Caryopsis in abaxial view. K. Caryopsis in abaxial view.