Fortipalpa pichincha, Supeleto & Santos & Aguiar, 2022

Supeleto, Fernanda A., Santos, Bernardo F. & Aguiar, Alexandre P., 2022, Revision of Fortipalpa Kasparyan & Ruíz-Cancino, (Ichneumonidae, Cryptinae), Zootaxa 5219 (6), pp. 501-533 : 518-521

Fortipalpa pichincha sp. nov.

( Figs 10–11 , 19–21 )

Description. Holotype ♀. Fore wing 9.10 mm. Head ( Figs 10 , 11A–B ). MLW 1.62. MWW 0.46. MSM 0.33. CHW 0.52. Clypeus anterior margin centrally with two small tubercles. Antenna with 23 flagellomeres. Supra-antennal area distinctly coriarious; median line distinct and straight.

Thorax ( Figs 10 , 11A ). Epomia approximately straight. Mesoscutum length to width 0.93 × as long as wide. Notaulus reaching about 0.39 of mesoscutum length. Scuto-scutellar groove with distinct longitudinal wrinkles. Epicnemial carina reaching about 0.8 of distance to subalar ridge; ending above pronotoal spiracle; almost straight. Sternaulus strong but narrow, posterior 0.3 nearly indistinct; weakly sinuous. Mesopleuron polished smooth, ventral half punctate. Mesosternum medio-longitudinal suture without transverse wrinkles. Transverse furrow anterior to propodeum centrally with very weak longitudinal wrinkles. Metapleuron densely punctate-rugulose; moderately pilose.

Propodeum ( Figs 10 , 11 A&D ). Area behind anterior transverse carina transversally strigate, arched posteriorwards. Spiracle elongate, SLW 2.06. Anterior transverse carina medially slightly arched forwards.

Wings (Figs X, Y). Fore wing crossvein 1m-cu more or less uniformly curved; fully continuous with 1-Rs+M. Vein 2Cua 0.76 × as long as crossvein 2cu-a. APH 0.67. Areolet subpentagonal; distinctly wider than high, AWH 0.71. Crossvein 3r-m spectral. Crossvein 2r-m and 3r-m about same length. Vein 3-M distinctly shorter than 2-M. HW1C 0.97. Hind wing vein Cub straight. Vein 2-1A ending at 0.13 of distance to wing margin.

Metasoma ( Figs 10 , 11A&D ). T1LW 2.46. T1WW 1.98. T1 spiracle at 0.57 of its length. Median depression of T1 elongate, distinct. T2LW 0.91. T2WW 1.92. T2–8 from distinctly coriarious T2 to smooth T8. Ovipositor slightly upcurved, OST 1.46; ventral valve apex with 13 teeth ( Fig. 11D ).

Color. Head pale yellow (231,209,132) with supra-antennal area and vertex black (36,31,21). Clypeus mid ventrally and supraclypeal area latero ventrally with black spot. Malar space black. Orbital band narrow, almost complete, briefly interrupted near 6 h. Mandible basal 0.25 pale yellow (231,209,132), otherwise black. Antenna black with short whitish band (233,215,149), composed by fl6 apical 0.30, fl7–10, and tiny dorsal spot basally on fl11. Pronotum black, collar light yellow (228,205,113). Propleuron light orange (222,132,54). Mesoscutum black, median lobe centrally with light yellow spot. Tegula and subalar ridge light yellow. Mesosoma otherwise mostly light orange. Scutellum light yellow, anteriorly black (37,25,8), posterior apex with narrow light orange stripe; yellow area somewhat bilobed. Postscutellum light orange. Fore leg: coxa, trochanter, trochantellus and femur light orange (218,149,59); tibia light yellow; t1–2 pale yellow (221,194,124); t3 pale yellow, apical 0.30 brownish (170,125,81); t4 brown (124,80,54); t5 dark brown (61,51,38). Mid leg: coxa, trochanter, trochantellus and femur light orange; tibia light yellow; t1–2 pale yellow, t2 apical 0.30 brown (124,80,54); t3–5 dark brown. Hind leg: coxa light orange (222,132,54); trochanter, trochantellus and femur fulvous (176,105,60); tibia bright yellow (209,160,35), apical 0.10 dark brown; t1 bicolored, basal 0.60 dark brown, apical 0.40 whitish (243,239,227); t2–4 whitish; t5 whitish, apically dark brown. Metasoma striped, dark brown (45,29,16) and yellow (231,209,132). T1 basal half dark orange (188,86,43), subapically black (36,31,21), posterior 0.2 pale yellow (231,209,132). T2–8 bicolored, dark brown with posterior 0.20–0.50 pale yellow. S2–4 anteriorly brown, posteriorly pale yellow.

Morphometry. See Table 1, Figs 20–21 , and Appendix.

Male. Unknown.

Comments. Structure. Most similar to F. panamensis , but differs by having the following features: Scutoscutellar groove with distinct longitudinal wrinkles (vs. with vestigial longitudinal wrinkles); sternaulus strong but narrow, posterior 0.3 nearly indistinct (vs. moderately strong, wide and shallow at posterior 0.25); T1LW 2.46 (vs. 2.67–2.85); HW1C 0.97 (vs. 1.30–1.49); vein 2-1A ending at 0.13 (vs. 0.20–0.26) of distance to wing margin; epomia after diverging from pronotal collar approximately straight (vs. distinctly curved); mesopleuron polished smooth, ventral half punctate (vs. polished smooth, centro-anteriorly with transversal striation, ventral half punctate); OST 1.46 (vs. 1.58–1.59); hind wing vein Cub straight (vs. distinctly convex on posterior half, otherwise straight); ovipositor ventral valve apex with 13 teeth (vs. 14–15); ovipositor slightly upcurved (vs. straight).

Also similar to F. frida , but differs by having the following features: Clypeus anterior margin centrally with two small tubercles (vs. without teeth); sternaulus weakly sinuous (vs. strongly sinuous); scuto-scutellar groove with distinct longitudinal wrinkles (vs. with vestigial longitudinal wrinkles; sternaulus strong but narrow, posterior 0.3 nearly indistinct (vs. moderately strong, wide and shallow at posterior 0.25); HW1C 0.97 (vs. 1.70–2.00); hind wing Vein 2-1A ending at 0.13 (vs. 0.20–0.27) of distance to wing margin; MWW 0.46 (vs. 0.51); T2LW 0.91 (vs. 1.01–1.08); notaulus reaching about 0.39 of mesoscutum length (vs. 0.45–0.49); mesosternum medio-longitudinal suture without transverse wrinkles (vs. with short transverse wrinkles); epicnemial carina reaching about 0.8 of distance to subalar ridge (vs. reaching about 0.7 of distance to subalar ridge); fore wing 9.10 mm (vs. 6.45–7.95 mm); ovipositor slightly upcurved (vs. straight).

Also similar to F. exelysae , from which it differs by having the following features: Area behind anterior transverse carina transversally strigate, arched posteriorwards (vs. transversally strigate, generally weak, anterolaterally faint); sternaulus strong but narrow, posterior 0.3 nearly indistinct (vs. moderately strong, wide and shallow at posterior 0.25); HW1C 0.97 (vs. 1.17–1.68); mesopleuron polished smooth, ventral half punctate (vs. polished smooth, centro-anteriorly with transversal striation, ventral half punctate); metapleuron densely punctate-rugulose (vs. densely punctate); T2–8 from distinctly coriarious T2 to smooth T8 (vs. faintly coriarious); vein 2-1A ending at 0.13 (vs. 0.17–0.26) of distance to wing margin; fore wing crossvein 1m-cu fully continuous with 1-Rs+M (vs. limit between 1m-cu and 1-Rs+M more or less traceable); hind wing vein Cub straight (vs. almost straight, very slightly convex); transverse furrow anterior to propodeum centrally with very weak longitudinal wrinkles (vs. entirely polished smooth); ovipositor slightly upcurved (vs. straight).

Color pattern. Closest to F.sacha , from which it differs by having pronotum collar yellow (vs. black); propleuron light orange (vs. dark brown); mandible apical 0.65 blackish (vs. apical 0.30 dark brown); and hind t5 whitish (vs. dark brown).

Also similar to F. exelysae , from which it differs by having dorsal margin of pronotum black (vs. yellow); epicnemium light orange (vs. mostly or almost entirely black); propodeum posteriorly without pale yellow mark (vs. with pale yellow mark); mandible apical 0.65 blackish (vs. apical 0.25 dark brown); hind t1 basally blackish (vs. yellowish); and scutellum antero centrally with black spot (vs. entirely pale yellow).

Etymology. The specific epithet is a literal reference to the municipality where the holotype was collected.

Distribution records. Ecuador ( Fig. 19 ).

Material examined. One specimen, 1♀. Holotype: ECUADOR, 1♀, Pichincha, Mindo , 2?, 1400 m, 0°03′04.29″S, 78°46′41.89″W, 2005-10, M.Cooper,?, FAS 4627 , Cooper coll. BNHH[E] 2005-152, NHM.

Figures

FIGURE 10. Fortipalpa pichincha sp. nov. Holotype female. Habitus.

FIGURE 11. Fortipalpa pichincha sp. nov. Holotype female. A, Dorsal view. B, Head, frontal. C, Propodeum, dorsal. D, Ovipositor tip, left side.

FIGURE 19. All known distribution records for Fortipalpa spp. (first map) and individual records for each species. Taxa presented in alphabetical order.

FIGURE 20. All results for morphometric ratios calculated for specimens of Fortipalpa spp., shown comparatively. On each plot, the species are sorted on the Y-axis from highest (top) to lowest (bottom) average. Graphs organized by alphabetical order of ratios.

FIGURE 21. Continuation of Figure 20.