Glaresis sabulosa Gordon and Hanley, 2014

Gordon, Robert D. & Hanley, Guy A., 2014, Systematic revision of American Glaresidae (Coleoptera: Scarabaeoidea), Insecta Mundi 2014 (333), pp. 1-91 : 24-26

Glaresis sabulosa Gordon and Hanley , new species

Description. Male. Length 4.0 mm, width 2.3 mm body form elongate, robust, widened from elytral base to apical 1/3 ( Fig. 15A ). Color yellowish brown. Head with clypeal surface finely rugose with small, sparse tubercles extended onto frons, frons finely rugose, vertex finely rugose without basal carina, setae short, indistinct. Clypeal apex slightly raised, weakly sinuate with median emargination, lateral angles angulate ( Fig. 15B ). Mandible pair symmetrical; mesal tooth strong; lateral prominence weak, outer margin abruptly rounded. Pronotum with distinct, long, transverse fovea in anterior 1/4 extended completely across pronotum, small slightly transverse fovea on each side of middle in anterior 1/2, wide, deep fovea on each side medially near lateral margin, distinct central furrow extended from base to transverse furrow; surface finely rugose, with sparse, straight setae-bearing carinae, setae decumbent, about 1/2 length of carina; anterior and lateral pronotal margins not bordered, all margins crenulate. Elytra with surface dull, finely microreticulate; all striae convex, weakly carinate, carinal segments small, widely separated, not much more than setal bases, seta much longer than base; intervals with large, shallow, slightly rectangular punctures ( Fig. 15C ). Metasternum long, dull, finely, densely microreticulate, median surface without ridges, laterally with scattered, straight ridges in basal 2/3, median area flat, with trace of median carina extended anteriorly from apical keel 1/2 distance to mesocoxae ( Fig. 15D ); metasternal groove shallow, clearly demarked. Lateral protibial teeth unevenly spaced, basal 2 teeth closer together. Mesotibia with 6 spines laterally, spinal bases close together, tibia slightly projecting at apex ( Fig. 15D ). Posterior metatrochanteral margin without small teeth; posterosuperior surface of metatrochanter with single tooth ( Fig. 15G ). Metafemoral surface with widely, evenly spaced, setaebearing tubercles, microreticulate; width to length ratio 1.0:1.5, without noticeable flange on anterior margin; posterior metafemoral margin angulate next to trochanter; posterosuperior margin with 2 widely spaced teeth, outer tooth small ( Fig. 15G ). Metatibia broadly triangular, surface entirely microreticulate, outer margin with series of small teeth from base to posteromedian lateral projection, projection small, oblique, surface smooth, median row of tubercles nearly lacking, inner margin smooth, pubescent ( Fig. 15F ). Apical margin of male 5th abdominal ventrite feebly rounded. Genitalia long, basal piece about as long as parameres, proximal end curved; median lobe shorter than parameres, tapered from base to apex, narrower at middle than a paramere, slightly curved upward in apical 1/3, apex rounded in ventral view; parameres slightly curved before apex in lateral view, margin straight, apex narrowly rounded ( Fig. 15E ).

Female. Apex of 5th abdominal ventrite strongly rounded.

Variation. Length 3.3 to 4.0 mm, width 1.8 to 2.3 mm. Number of lateral mesotibial spines varies from 6-8; number of tubercles and form of lateral projection on metatibia vary slightly, number of teeth on posterosuperior metafemoral surface varies from 1 to 2; posterosuperior teeth on metatrochanter vary from 1 to 2 closely spaced teeth.

Type material. Holotype male: Utah: UTAH: (Garfield Co.) 26 mi. S. Hanksville , Hwy. 95, 16-VI-84, Robert Gordon ( USNM) . Paratypes, 34: (15), same data as holotype; (1) Utah, Emery Co., Little Gilson Butte , 29-VIII-1986, A. S. Menke; (1) Wayne Co., Hanksville , 28-V-1974; (1) 14 mi. S. Hanksville, Fairview Rch., 14-VI-82, Robert Gordon; (6) Emery Co., Goblin Valley , 1-VI-82; (3) Utah, Emery Co., Goblin Valley Rd. at Hwy 24 ; UV trap; M. Raschko; 16-17.V.2007; (1) Utah, Emery Co., Wildhorse Cr N of GoblinVly 4900', 27-28 May 1985, F.D. Parker Collector, D.K. Broemeling Collector; (9) Emery Co., 17 mi. N. Hanksville, “dunes” nr. Gilson Butte well, VII-27-78, Andrews & Hardy ;1); Utah, Garfield Co., 42 km SSE Hanksville , 15 June 1982, W. Steiner, R. Gordon & D. Whitehead, colls.; ( CASC) ( CSCA) ( MJPC) ( USNM) ( USUL) .

Other specimens. 276: Arizona: Navajo Co., Betatakin Ruins. Colorado: Maybell. Utah: (71) Kane Co. , Coral Pink Dunes , 10-VII-1980, Robert Gordon ; same data except collection date 18-VI-1993; (3) Kane Co., Coral Pink Dunes , 16 km W Kanab, 2045m, 26 Aug.-7 Sep. 1991, J. M. Hill, C. B. & R. Knisley, W. E. Steiner collectors; (21) Kane Co., Coral Pink Sand Dunes, BLM land NW Kanab , 37 o 05’14". 112 o 40’11", P. Skelley and family; (22) Kane Co., Coral Pink Sand Dunes State Pk , 14-VII-1998, Baumann, et al.; (35) Kane Co., Coral Pink sand dunes, June 4, 1981, Baumann; (5) Kane Co., Coral Pink Sand Dunes St. Pk. , VIII-2/3-85, Rolf L. Aalbu col. BL; (3) Kane Co., Coral Pink Sand Dunes, 1.8 mi. S. Hancock Rd, on Sand Dunes Rd. ; (7) Kane Co., Ponderosa Cmpgd , Coral Pink Sand Dunes; (4) Kanab Co., 8 mi. N. Kanab , 29 Aug 1962, G. F. Knowlton; (43) Garfield Co., 7.9 mi. SE Boulder, Sand Dunes , 37 o 52’18" 111 o 20'’14"W, 25-VII-2001, RD Gordon; (1) Garfield Co., Grand Staircase-Escalante Natl. Mon., Sand Creek Boulder Mail Trail, 31 July-1 August 2001; (1) Garfield Co., Burr Trail between Deer Creek and Sleep Creek , 23-VII-2001, R.W. Baumann; (1) Washington Co., Sand Hollow Rd., E. entr. St. Pk. , UV light, W.B. Warner; (2) Utah, Washington Co., Zion Natl’l Park , 8 Aug 1966, R. Peckenpaugh ; Washington Co., 6 mi. SW Hurricane sand dunes. Wyoming: (28) Sweetwater Co., Flaming Gorge , Blacks Fork River , 14-VI-1993, Col: R. Gordon, UV light; (6) Sweetwater Co., Flaming Gorge Res., BF. & J. Carr, pitfall trap, ant mound; (42) Sweetwater Co., Red Desert , sand dunes, 23-VI-1977, Gordon & Ashworth; (1) Sweetwater Co., Hwy 191, 7 mi. S I-80, 14-VI-84, Robert Gordon. ( BYUC) ( CMNH) ( FSCA) ( JCIC) ( MJPC) ( RCCC) ( USNM) ( USUL) .

Remarks. Glaresis sabulosa is known from several southern Utah localities, but specimens from each locality vary slightly from the typical and from each other. Therefore the type series is restricted to sand deposits in the vicinity of Hanksville in Emery, eastern Garfield, and Wayne Counties. Specimens from Kane Co., Coral Pink dunes, differ from the typical by somewhat larger tubercles on clypeus and frons, and by distinctly more tubercles and more robust posteromedian projection of the metatibia. Specimens from dunes southeast of Boulder also have more prominent head tubercles, and the metatibial sculpture in some (not all) specimens is more pronounced than in Coral Pink specimens. However, all specimens examined have the shallow but distinctly defined metasternal groove that helps define G. sabulosa . Southern Wyoming specimens are distinctly larger and differ by having more prominent head tubercles, darker color, and slightly different metatibial posteromedian projection. The Wyoming specimens are perhaps another species, but we are unable to satisfactorily distinguish them from Utah G. sabulosa on morphological characters.

Glaresis inducta is the most similar species, but has 4 or 5 metatibial spines; lacks a definable metasternal groove; body short, robust; and elytral carinae nearly continous.

All type specimens have been taken at light, and specimens collected by one of us (RDG) in the Hanksville area were at ultra violet light in blow sand deposits such as dunes and sand fields, etc. Nontype specimens were also taken in areas of sand deposits, all Wyoming specimens were collected at UV light (R. Gordon, pers. obs.)

Etymology. The specific epithet is the Latin sabulosus, meaning sandy, referring to the extremely sandy habitat of this species.

Figures

Figures 15A-15G. Glaresis sabulosa, new species. A) Habitus view. B) Clypeus and head surface. C) Pronotal surface. D) Metasternum and mesotibia. E) Male genitalia. F) Metaleg and trochanter, dorsal view. G) Metaleg and trochanter, ventral view.