Glyphiulus mediobliteratus, Golovatch & Geoffroy & Mauriès & Spiegel, 2007

Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul & Spiegel, Didier Van Den, 2007, Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 2: the javanicus-group, Zoosystema 29 (3), pp. 417-456 : 426-430

publication ID

https://doi.org/ 10.5281/zenodo.5393630

persistent identifier

https://treatment.plazi.org/id/B36487D8-FFC8-D535-D0F2-FA5BFB03FA9E

treatment provided by

Marcus

scientific name

Glyphiulus mediobliteratus
status

sp. nov.

Glyphiulus mediobliteratus View in CoL n. sp. ( Figs 6-8 View FIG View FIG View FIG )

TYPE MATERIAL. — Vietnam. Ninh Binh Prov., Cuc Phuong National Park, Cave 4: Song Cung Cave , 21. VI.2003, leg. P. Trontelj & B. Sket, holotype ♂ ( MNHN GA 046 About MNHN ) ; paratypes 3 ♂♂, 2 ♀♀, 1 fragm., 1 juv. ( MNHN GA 046 About MNHN ) ; 1 ♂ ( SEM) ; 1 ♀ ( ZMUC) ; 1 ♀ ( ZMUM) ; 1 ♀ ( MCSNV) ; 1 ♀ ( NMNHS) ; 1 ♀ ( OBBFUL). — Same locality, Cave 2: Thanh Yen Water Cave , 20. VI.2003, leg. P. Trontelj & B. Sket, paratypes 1 ♂ ( ZMUM) ; 1 ♀ ( MNHN GA 046 About MNHN ) .

ETYMOLOGY. — To emphasize the strongly obliterated collum and axial crests.

DIAGNOSIS. — Differs from congeners by the unpigmented ocelli and, in part, body, coupled with the nearly smooth collum, the rudimentary axial crests, the lack of a notch on each anterior gonopod half, the highly conspicuous, slightly plumose flagellum of the posterior gonopod, etc. (see also key below).

DESCRIPTION

Holotype with 61p+1a+ T, about 43 mm long and 2.0 mm wide; adult paratypes (♂, ♀) with 51-62p+2- 1a+ T, length 32-42 mm, width 1.8-2.0 mm.

Coloration entirely pallid to pale brown, only tips of tubercles of tergal crests often brownish to dark brown. Ocelli not visible.

Antennae ( Figs 6A View FIG ; 8A View FIG ), gnathochilarium (mentum divided, n = 5) ( Fig. 8B View FIG ), segment shape ( Fig. 7A View FIG ), epiproct ( Fig. 6E View FIG ), paraprocts, hypoproct, male legs 2 and 3 ( Fig. 8D, E View FIG ), texture ( Fig. 6 View FIG A- E) generally as in G. obliteratus n. sp., but sparse striation present in front of stricture both laterally and dorsally; conspicuous pleurosternal crests on segments 2-4 wanting ( Fig. 6A View FIG ); collum ( Fig. 6A, B View FIG ) with 1a-4a+ma+4a-1a very weak undulations in caudal 1/3, undulation ma being shortest; tergal crests higher, even adding another lateral crest below ozoporiferous one; ozoporiferous cones especially high, about as high as wide ( Fig. 7A View FIG ), each crest

Golovatch S. I. et al.

distinctly subdivided into two usually subtriangular halves, except for nearly absent axial crests (m); carinotaxy pattern thus 2/2+I/i+1/1+m+1/1+I/ i+2/2 ( Figs 6 View FIG A-E; 7A); a small accessory spine present at base of claw on legs 2 and 3 only ( Fig. 8D, E View FIG ), thereafter virtually missing ( Fig. 7F View FIG , 8F View FIG ); legs slightly shorter than body diameter ( Fig. 7A View FIG ); male legs 1 with strongly developed paramedian coxal outgrowths and 5-segmented, slightly reduced, telopodites ( Fig. 8C View FIG ).

Head width = segment 2 = 6 <collum = 7(8)> segment 3 = 5 <7 <8 <9 <10 <11 = midbody segments; body gradually but then abruptly tapering toward telson. Postcollar constriction modest ( Fig. 6B View FIG ).

Penes broad, rounded, each with three or four strong setae distolaterally ( Fig. 8D View FIG ).

Anterior gonopods ( Figs 7 View FIG B-D; 8G) without notches, processes (d) especially high. Posterior gonopods ( Figs 7E View FIG ; 8H View FIG ) with characteristic, long, distally plumose flagella.

REMARKS

The presence of completely unpigmented ocelli suggests that this species may be troglobitic, but further observations are needed to confirm this.

VI

Mykotektet, National Veterinary Institute

MNHN

Museum National d'Histoire Naturelle

ZMUC

Zoological Museum, University of Copenhagen

ZMUM

Zoological Museum, University of Amoy

T

Tavera, Department of Geology and Geophysics

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