Goeldia guayaquilensis ( Schmidt, 1971 )

Almeida-Silva, Lina M. & Brescovit, Antonio D., 2024, Unraveling the mysteries of Goeldia Keyserling, 1891: revision, description of seven new species and first record from USA (Araneae: Titanoecidae), Zootaxa 5428 (2), pp. 151-193 : 170-171

Goeldia guayaquilensis ( Schmidt, 1971)

( Figs 13 ; 25 )

Titanoeca guayaquilensis Schmidt, 1971: 418 , fig. 23 (holotype male, collected in Germany but coming in a banana shipment from Ecuador (2°0’0”S 77°30’0”W), deposited in FMS 25791/1, examined).

Goeldia guayaquilensis : Dupérré 2023: 271, figs. 100a–b, 101a–b; World Spider Catalog 2023.

Additional Material Examined. ECUADOR. Santo Domingo de los Tsáchilas: near Santo Domingo de Los Colorados, Tinalandia (0°19’12”S 78°58’49”W), 3♀, 7.XII.1981, E. Schlinger coll. ( CAS 9025939 ) . Pichincha: Guayllabamba (0°3’33”N 78°20’29”W), 30 Km E of Quito, before the river of Guayllabamba , 12♀, 24.VIII.1978, A. Roig coll. ( MACN 16059 ) ; Valle Guayllabamba (0°3’32.59”S 78°20’29.21”W), 10♀, 7.V.1982, A. Roig coll. ( MACN16054 ) . Tungurahua: Baños (1°23’47”S, 78°25’29”W), 1♀, 24.III.1984, A.Roig coll.( MACN16062 ) ; Baños (1°23’47”S, 78°25’29”W), 1♀, IX.1978, A. Roig coll. ( MACN 16039 ) . Morona-Santiago: Near Macas (2°19’75”S 78°08’2.88”W), 24.III.2011, A. Chagas et al. coll. ( MNRJ 6388 ) . Azuay: Gualaceo (2°54’0”S 78°47’0”W), 2♀, 1♂, 1.IX.1978, A. Roig coll. ( MACN 16123 ) . Loja: Vilcabamba (4°15’21”S 79°13’8”W), 12 Km S Loja, 1♀, 12.VII.1978, A. Roig coll. ( MACN 16041 ) . PERU. Cuzco: Urubamba, Ollantaytambo (13°15’31”S 72°15’49”W), 3♀, 13.I.1983, A. Roig coll. ( MACN 16124 ) . Lima: Cañete, Valle de Mala (12°39’27”S 76°37’45”W), Santa Enriqueta , 1♀, 19.IX.1997, M. A. Benamu-Pinto coll. ( IBSP 14902 ) ; near Lima (12°2’6”S 77°1’7”W), Lake Villa , 1♂, X.1951, V. Weyrauch coll. ( CAS 9025991 ) ; Barranca, 36.8 Km N Pativilca (10°41’4”S 77°46’43”W), 1♀, 1 imm., 15.I.1953, E. I. Schlinger & E. S. Ross coll. ( CAS 9025983 ) .

Diagnosis. Males of G. guayaquilensis differ from others species by the reduced and narrow tegular process ( Fig. 13A–B ) and by the blunt and dorsally projected patellar apophysis ( Fig. 13B–D ) in the male palp. Female differs by the epigynum with posterior border straight ( Fig. 13E ), thin spermathecae when compared to the copulatory ducts ( Fig. 13F ), and vertical spermathecae short, with half of the height of the copulatory ducts ( Fig. 13F ).

Description. Male (Holotype, FMS 25791/1). Coloration: carapace, labium, endites and sternum yellowish brown; chelicerae reddish brown; legs I–III with femora brown; leg IV brown from femur to metatarsus; other leg segments yellowish brown. Total length: 3.38. Carapace: length 1.88, width 1.34, height 0.86. Opisthosoma: length 1.50, width 1.00. Clypeus: height 0.07. Eye diameters and interdistances: AME 0.05; ALE 0.09; PME 0.07; PLE 0.07; AME–AME 0.08; AME–ALE 0.17; ALE–PLE 0.05; PLE–PME 0.15; PME–PME 0.12; PME–AME 0.09. Paturon: 0.98 long. Leg measurements: I: femur 1.75/ patella 0.55/ tibia 1.65/ metatarsus 1.48/ tarsus 0.8/ 6.23; II: 1.59/ 0.62/ 1.5/ 1.38/ 0.75/ 5.84; III: 1.5/ 0.5/ missing/ missing/ missing; IV: 1.8/ 0.58/ 1.74/ 1.23/ 0.65/ 6.0. Leg formula: 1423. Leg spination: Femur I p1ap; II p1ap; Tibia II v0-2-0 or v0-1r-2-1r, r0-1-0-1, p1sa or p2sa; IV v0- 0-2ap; Metatarsus I v2-1r- 1m; II v2-1p-0 or v2-2- 1m, r0-0-1ap, p0-0-2; IV v2-2- 1m. Palp with patellar apophysis with same thickness along its entire length ( Fig. 13C ). Retrolateral side of the tibia strongly rounded and projected ( Fig. 13D ). Tegular process small, not reaching the edge of the tegulum ( Fig. 13B ).

Female (CAS 9025939). Coloration: carapace brown with cephalic area slightly darker; chelicerae, endites, labium brown, slightly darker than carapace; femora brown and other leg segments yellowish brown. Opisthosoma grayish brown. Total length: 3.80. Carapace: length 1.52, width 1.09, height 0.5. Opisthosoma: length 2.23, width 1.44. Clypeus: height 0.14. Eye diameters and interdistances: AME 0.06; ALE 0.08; PME 0.09; PLE 0.09; AME– AME 0.09; AME–ALE 0.06; PME–PME 0.08; PME–PLE 0.09; AME–PME 0.06; ALE–PLE 0.04. Paturon: 0.89 long. Leg measurements: I: femur 1.3/ patella 0.60/ tibia 1.13/ metatarsus 0.97/ tarsus 0.49/ total 4.49; II: 1.17/ 0.50/ 0.92/ 0.79/ 0.44/ 3.82; III: 1.06/ 0.46/ 0.69/ 0.74/ 0.35/ 3.3; IV: 1.35/ 0.48/ 1.1/ 0.86/ 0.44/ 4.23. Leg formula 1423. Leg spination: Femur I p1ap; Metatarsus I v1map, p1ap, r1ap; III v2ap, r1ap, p1ap; IV v1map, r1ap, p1ap. Metatarsus with more setae than other leg segments. Epigynum with posterior edge of the median field straight ( Fig. 13E ); copulatory ducts visible at the copulatory opening ( Fig. 13E ). Apex of the copulatory ducts dorsally projected, base enlarged ( Fig. 13F ). Horizontal and vertical spermathecae slim ( Fig. 13F ).

Variation. Length (2♂): total 3.35–3.38; carapace 1.87–1.88; femur I 1.75–1.76; (10♀): total 3.9–5.6; carapace 1.55–2.48; femur I 1.03–1.45.

Distribution. Peru and Ecuador ( Fig. 25 ).

Figures

FIGURE 13. A–F. Goeldia guayaquilensis (Schmidt, 1971). A–D: male, left palp. E–F: female epigynum: A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum, ventral view; F. Vulva, dorsal view.

FIGURE 25. Distribution of Goedia spp. in northern South America: Goeldia diva (pentagon—pink); G. obscura (square— yellow); G. guayaquilensis (circle—purple); G. mirim (star—red); G.utcuyacu (rectangle—blue). Type species are marked as black versions of each symbol when there are records to multiple localities.