Goja caldasensis, Carrejo & Obando & Aldrete & Mendivil, 2021

Carrejo, Nancy, Obando, Ranulfo González, Aldrete, Alfonso Neri García & Mendivil, Julián, 2021, New species of Goja Navás (Psocodea: ‘ Psocoptera’: Epipsocidae) from Colombia Mexico and Peru, Zootaxa 4903 (4), pp. 501-541 : 505

Goja caldasensis n. sp. Male.

( Figs 1–6 )

Diagnosis. Forewing Rs dichotomously branched, R2+3 forked, R4+5 dichotomously branched, resulting in four veins. M with five primary branches, M2, M3 and M5 forked as illustrated, resulting in ten veins. Hindwing Rs forked, M three branched, with M2 forked.Anterior pair of endophallic sclerites directed posteriorly, curved, distally acuminate; mesal pair with 2–3 long, acuminate processes directed outward ( Fig. 6 ). It is similar to G. pillcopatensis n. sp. and G. yarumosensis n. sp., differing from them in wing venation, phallosome and endophallic sclerites.

Color (in 80% ethanol). Body brown-yellowish. Head pale brown ( Fig. 3 ), postclypeus with ochre striae. Compound eyes dark brown, ocelli hyaline, with ochre centripetal crescents. Maxillary palps pale brown. Antennae pale brown. Thorax pale brown, pronotum cream, with brown spots; tergal lobes of meso- and metathorax with brown spots, mesopleura pale brown. Legs pale brown. Forewings ( Fig. 1 ), pale yellowish, veins pale brown. Hindwings ( Fig. 2 ), hyaline, with a yellowish hue, veins pale brown. Abdomen pale brown. Hypandrium brown, darker distally on sides; epiproct and paraprocts pale brown. Phallosome brown, with endophallic sclerites and aedeagal arch pale brown.

Morphology. As in diagnosis, plus the following: Head: H/MxW: 1.41; compound eyes large, H/d: 4.22; IO/ MxW: 0.74. Outer cusp of lacinial tips broad, with nine denticles. Mx4/Mx2: 1.08. Forewings ( Fig. 1 ): FW/W: 2.51; M 5-branched, resulting in 10 veins at wing margin as illustrated. Pterostigma: lp/wp: 3.46, areola postica tall, triangular: al/ah: 2.54. R2+3 forked, R4+5 dichotomously branched, resulting in four veins at wing margin; Hindwings ( Fig. 2 ): HW/w: 2.95. Rs two-branched, M three-branched, M2 forked. Hypandrium ( Fig. 5 ), concave posteriorly in the middle, setose as illustrated; phallosome ( Fig. 6 ) with external parameres dilated distally, apically truncate, bearing pores. Endophallic sclerites ( Fig. 6 ). Paraprocts ( Fig. 4 ), broadly triangular, with a macrosetae near sensory field, and a distal setal field, one setae distally spatulate; sensory fields with 28 trichobothria on basal rosettes. Epiproct ( Fig. 4 ) wide, semioval, rounded posteriorly, with two lateral macrosetae, setal field on distal fourth as illustrated, one macrosetae mesally.

Measurements. FW: 3737.5, HW: 2875, F: 950, T: 1560, t1: 710, t2: 240, ctt1: 26, f1: 700, f2: 550, Mx4: 260, IO: 400, D: 243, d: 180, IO/d: 2.22, PO: 0.74.

Specimen studied. Holotype male. COLOMBIA. Caldas, Manizales, Río Blanco Natural Reserve 5º4’24.3”N: 75º26’16.2”W, 2592 m., 15.I.2011. R. González & A. N. García Aldrete. Beating branches of vegetation.

Etymology. The specific epithet refers to Caldas, the Colombian Department where the holotype was found.

Figures

FIGURES 1‒6. G. caldasensis n. sp. Male. 1. Forewing. 2. Hindwing. 3. Front view of head. 4. Left paraproct and epiproct. 5. Hypandrium. 6. Phallosome. Scales in mm.