Goloboffia pachelbeli, Ferretti & Ríos-Tamayo & Goloboff, 2019

Ferretti, Nelson E., Ríos-Tamayo, Duniesky & Goloboff, Pablo A., 2019, Four new species of Goloboffia (Mygalomorphae: Migidae) from Chile, Zootaxa 4712 (2), pp. 251-268 : 260-264

Goloboffia pachelbeli sp. nov.

( Figs 48–71 )

Type material. Holotype: ♂, MACN-Ar 36292, Chile, Región de Coquimbo (IV), Elqui, Totoralillo Norte (29°33′33.1″S, 71°18′57.8″W), collected under stone, 12.vii.2009, M. Izquierdo, A. Ojanguren, J. Pizarro and F. Alfaro, leg .. Paratypes: 1 ♂, MACN-Ar 36292, same collection data ; 1 ♀, MACN-Ar 36123, Chile, Región de Co- quimbo (IV), Route 5, 35 km north of La Serena, 2.xi.1988, P. Goloboff leg.

Additional material examined. 1 juvenile, MACN-Ar 36123, Chile, Región de Coquimbo (IV), Route 5, 35 km north of La Serena , 2.xi.1988, P. Goloboff leg.

Etymology. The specific epithet is a patronym in honor of Johannes Pachelbel (1653–1706), a German composer and organist, and one of the most important composers of the middle Baroque era.

Diagnosis. Males of Goloboffia pachelbeli sp. nov shares with G. vellardi the general aspect of the palpal bulb, but differ from them by a shorter, thicker and uniformly curved embolus ( Figs 56–57 ) and by a more spinose metatarsi IV ( Fig. 54 ); females differ from the other known species of Goloboffia by the ocular group occupying almost the entire cephalic width (0.93) ( Fig. 64 ), in combination with the absence of spines on tarsi III and IV ( Figs 66–67 ). Females also differ by the shape of the spermathecae with long straight ducts 1.5 longer than spermathecal head width, large oval lobes ( Fig. 70 ).

Description. Holotype male, MACN-Ar 36292. Coloration (in alcohol): Carapace and chelicerae reddish brown, legs brown, abdomen brownish with grayish central chevron ( Figs. 48–51 ). Total length 9.04; carapace 4.73 long, 4.44 wide. Fovea deep, slightly recurved with recurved ends, 0.77 wide. Ocular group 1.89 wide, occupying 0.47 of cephalic width ( Fig. 50 ). Eye sizes and interdistances: AME 0.19, PME 0.14, ALE 0.39, PLE 0.13, AME–AME 0.15, AME–ALE 0.41, PLE–PME 0.11, PME–PME 0.77. Labium 0.66 long, 0.63 wide, without cuspules. Labium separated from sternum by shallow groove. Maxillae without cuspules ( Fig. 52 ). Fangs keeled and chelicerae with subequal teeth, 4 promarginal and 4 retromarginal, and 18 denticles between margins. Sternum elongated, 2.54 long, 2.08 wide. Posterior sigillae shallow and oval ( Fig. 49 ). Leg formula 4123. Segment lengths (femur, patella, tibia, metatarsus, tarsus, total): I: 4.25, 1.68, 2.94, 2.59, 0.84, 12.3. II:3.8, 1.65, 2.66, 1.89, 0.8, 10.8. III: 2.71, 1.29, 2.36, 1.69, 0.87, 8.92. IV: 3.81, 1.98, 3.06, 2.54, 1.14, 12.53. Palp: 2.67, 0.8, 2.46, –, 0.55, 6.48. Spines: Femur: Palp: 1d; I: 1 – 1 – 1d, 1 – 1 – 1 – 1 – 1 – 2p; II: 1 – 1 – 1d, 1 – 1p; III: 1 – 1d, 1r; IV: 1 – 1 – 1d. Patella: Palp: 0; I: 2p, 1 – 2v; II: 1p, 1v; III: 1 – 1 – 2d, 1r; IV: 0. Tibia: Palp: 0 ( Figs. 58–59 ); I: 1 – 1 – 1 – 1p, 2 – 1 – 1 – 2 – 1 – 2v ( Fig. 53 ); II: 2 – 2 – 1 – 2 – 2v, 1 – 1 – 1p; III: 1 – 2v; IV: 2v. Metatarsus: I: 1 – 1 – 1p, 1 – 1 – 1r; II: 1 – 1 – 1 – 1p, 1 – 1 – 1 – 1r; III: 1 – 1p ( Fig. 55 ); IV: 1 – 2v, 1p, 1r ( Fig. 54 ). Tarsus: I – IV: 0.

Bulb globose on base with short and thick embolus uniformly curved and expanded at tip ( Figs. 56–59 ).

Female (MACN-Ar 36123), coloration (in alcohol): Carapace, legs and chelicerae brown, abdomen light gray ( Figs 60–62 ). Total length 12.25; carapace 5.09 long, 4.23 wide. Fovea deep, straight with recurved ends, 1.14 wide. Ocular group 3.31 wide, occupying 0.93 of cephalic width ( Fig. 64 ). Eye sizes and interdistances: AME 0.79, PME 0.84, ALE 2.54, PLE 0.63, AME–AME 0.8, AME–ALE 3.57, PLE–PME 0.66, PME–PME 4.99. Labium 0.86 long, 0.85 wide, with 6 cuspules. Labium separated from sternum by shallow groove. Maxillae with 10/7 (right/left) cuspules ( Fig. 65 ). Fangs keeled and chelicerae with subequal teeth, 4 promarginal and 4 retromarginal, and 9 denticles between margins. Sternum elongated 3.03 long, 2.31 wide. Posterior sigillae shallow, small and airfoil shaped ( Fig. 63 ). Leg formula 4123, tibiae and metatarsi I and II flattened, with very long spines ( Figs 68–69 ). Segment lengths (femur, patella, tibia, metatarsus, tarsus, total): I: 3.5, 1.97, 2.1, 1.6, 0.79, 9.96. II: 2.72, 1.42, 1.65, 1.15, 0.54, 7.48. III: 2.28, 1.45, 1.41, 1.3, 0.81, 7.25. IV: 3.52, 2.04, 2.52, 2.67, 0.95, 11.7. Palp: 1.9, 0.82, 0.91, –, 1.11, 4.74. Spines: Femur: Palp and legs I – IV, 0. Patella: Palp: 1p; I: 1p; II: 1p; III: 15p; IV: 45d. Tibia: Palp: 1p, 1r; I: 6p, 11r; II: 5p, 5r; III: 13p; IV: 0. Metatarsus: I: 5p, 6r; II: 5p, 5r; III: ventral comb of 7 large and 14 small spines ( Figs 66–67 ); IV: ventral comb of 13 large spines. Tarsus: Palp: 3p, 3r, 6v; I: 3p, 4r, 8v; II: 3p, 8r, 10v; III: 0; IV: 0.

Spermathecae having oval lobes that point outwards with a well-developed constriction between lobes and ducts only in outer faces of receptacles, ducts very long (1.5 longer than spermathecal head width) ( Fig. 70 ).

Distribution. Known from Totoralillo and La Serena at Elqui province ( Fig. 71 ).

Figures

FIGURE 71. Known distribution of Goloboffia species in Chile.

FIGURES 48–55. Goloboffia pachebeli sp. nov., holotype male, MACN-Ar 36292. 48. Carapace, dorsal view. 49. Labium and sternum. 50. Eye group. 51. Abdomen, dorsal view. 52. Labium and maxillae. 53. Tibia I, ventral view. 54. Metatarsus and tarsus IV, ventral view. 55. Metatarsus and tarsus III, ventral view. Scale bars=1 mm.

FIGURES 56–59. Goloboffia pachebeli sp. nov., holotype male, MACN-Ar 36292. 56. Palpal bulb, dorsal view. 57. Palpal bulb, ventral view. 58. Palp, retrolateral view. 59. Palp, prolateral view. Scale bars=1 mm.

FIGURES 60–65. Goloboffia pachebeli sp. nov., paratype female, MACN-Ar 36123. 60. Carapace, dorsal view. 61. Carapace, lateral view. 62. Abdomen, ventral view. 63. Labium and sternum. 64. Eye group. 65. Labium and maxillae. Scale bars=1 mm.

FIGURES 66–70. Goloboffia pachebeli sp. nov., paratype female, MACN-Ar 36123. 66. Tarsus III, ventral view. 67. Tarsus III, retrolateral view. 68. Metatarsus I, prolateral view. 69. Tibia I, prolateral view. 70. Spermathecae, dorsal view. Scale bars=1 mm.