Gonaphodiellus pacatus ( Harold, 1880 )

Dellacasa, Marco, Dellacasa, Giovanni & Gordon, Robert D., 2012, Systematic revision of Gonaphodiellus taxa, with description of two new genera and fourteen new species (Coleoptera: Scarabaeidae: Aphodiinae), Insecta Mundi 2012 (230), pp. 1-41 : 12-13

Gonaphodiellus pacatus ( Harold, 1880)

( Fig. 51-56 )

Aphodius pacatus Harold, 1880: 37 ; Blackwelder 1944: 213.

Aphodius (Gonaphodiellus) pacatus ; Schmidt 1913: 133; Dellacasa 1988: 174.

Gonaphodiellus pacatus ; Paulian 1942: 73; Dellacasa et al. 2001: 155; Skelley et al. 2007: 4.

Type locality. Fusagasugà ( Colombia).

Type repository. Muséum national d’Histoire naturelle, Paris, France (type examined).

Redescription. Length 5.5-6.0 mm, oval-elongate, feebly convex; moderately shiny, glabrous. Brownyellowish, epistome, pronotal and elytral disc usually darker; legs brown-yellowish; antennal club pale testaceous. Head with epistome slightly convex on disc, finely, rather sparsely and subregularly punctured; clypeus faintly sinuate at middle, round at sides, very finely bordered, edge glabrous, lateral margins feebly arcuate; genae obtusely round, very shortly ciliate, protruding from the eyes; frontal suture finely impressed, not tuberculate, front finely, rather sparsely, subregularly punctured. Pronotum transverse, moderately convex, subparallel-sided, dually, somewhat irregularly punctured; punctation coarser and denser on sides; large punctures, twice to three times larger than small ones, almost lacking on disc, small punctures regularly scattered throughout, more superficial on disc; lateral margins rather thickly bordered, edge glabrous; hind angles widely truncate, truncature distinctly inwardly sinuate, base bisinuate. Scutellum elongate, finely sparsely punctured on basal half. Elytra slightly broadened posteriorly, moderately convex, weakly denticulate at shoulder; striae fine, superficially punctured, subcrenulate; interstriae feebly convex, sparsely superficially punctured. Male: pronotum not narrowed anteriorly; fore tibiae relatively slender and more elongate, their spur stouter and downward bent; metasternal plate longitudinally excavate; aedeagus Fig. 55-56 . Female: pronotum narrowed anteriorly, relatively more densely and coarsely punctured on disc; fore tibiae shorter and stouter, their spur slender and almost straight; metasternal plate nearly flat.

Material examined. COLOMBIA: Fusagasugà (1 ex., female, type, MNHN) ; Rovira , Tolima, VIII.1980, leg. L. Peña (1 ex., DCGI) ; COSTA RICA: Monte Verde , Puntarenas, V.1979, leg. Forsyth A. (1 ex., WBC) ; 4 mi. E Volcán , 09.XII.1952 (1 ex., USNM) ; PANAMA: Chiriqui Prov., Cerro Pelota , 4 Km N Sta. Clara, m 1500, VIII.1982, leg. Gill B., flight interception trap (1 ex., CNCI) ; Chiriqui Prov., Cerro Punta , m 1500, 23.V.1977, leg. Howden H., cup traps (12 exx., CNCI) ; idem, 5100’, 08.I.1964, leg. Bottimer L. J., horse dung (3 exx., CNCI) ; Chiriqui Prov., 2 Km W Cerro Punta, 8°51’N – 82°36’W, 22.V.1987, leg. Howden H., cup traps (4 exx., CMNO) ; idem, m 1720, 01-07.VI.1977, leg. Howden H. & A. (5 exx., CMNO) ; Chiriqui [Prov.], Hartmann’s Finca , 19.V.1996, leg. Turnbow R., feces baited pitfall (1 ex., RHTC) ; idem, 19-20.V.1995, leg. Turnbow R., feces baited pitfall (8 exx., DCGI, RHTC) ; idem, 04-07.VII.1997, leg. Morris & Wappes, feces baited pitfall traps (23 exx., DCGI, MCGA) ; idem, 18-20.V.1996, leg. Wappes, Huether & Morris (6 exx., FSCA) ; Chiriqui [Prov.], Sta. Clara, Hartmann’s Finca , 23-25.V.1980, leg. Riley E. & LeDoux (1 ex., DCGI) .

Distribution. Colombia, Costa Rica, Panama.

Bionomics. Poorly known; probably coprophagous species collected mostly in spring and often at light.

Figures

Figure 43-56. Gonaphodiellus opisthius (Bates, 1887) (Zacualtipan, Hidalgo, Mexico). 43) Epipharynx. 44- 47) Habitus (length ideogram, color variations and morphological details). 48-49) Aedeagus (dorsal and lateral view). 50) Right labial palp (dorsal view). Gonaphodiellus pacatus (Harold, 1880) (Tolima, Rovira, Colombia). 51) Labial palp. 52) Epipharynx. 53-54) Habitus (morphological details and length ideogram). 55-56) Aedeagus (dorsal and lateral view).