Gymnanthemum extensum (DC.) Steetz, Naturw. Reise Mossambique [Peters] 6(Bot., 2): 337. 1864.

Bunwong, Sukhonthip, Chantaranothai, Pranom & Keeley, Sterling C., 2014, Revisions and key to the Vernonieae (Compositae) of Thailand, PhytoKeys 37, pp. 25-101 : 72-73

Gymnanthemum extensum (DC.) Steetz, Naturw. Reise Mossambique [Peters] 6(Bot., 2): 337. 1864. Figs 9A-B

Vernonia extensa DC., Prodr. 5: 33. 1836.

Conyza extensa Wall., Numer. List [Wallich] no. 3061, comp. no. 126, nom. nud.

Gymnanthemum extensum Type. Nepal; Wallich 3061 (holotype: G!).

Vernonia cylindriceps C.B. Clarke, J. Linn. Soc. Bot. 25: 35. 1890.

Gymnanthemum cylindriceps (C.B.Clarke) H.Rob., Proc. Biol. Soc. Washington 112(1): 241. 1999.

Gymnanthemum extensum Type: India, Hartook Mekong, C.B. Clarke 42109 (holotype: K!).

Description.

Shrubs or subshrubs 2-6 m tall. Stems caulescent, young branches inconspicuously ribbed, white puberulous. Leaves 7-13 by 2-4 cm, oblanceolate, margin serrate, apex acute, base attenuate, chartaceous; upper surface puberulous, without glands; lower surface puberulous with whip-shaped hairs, cylindrical hairs and capitate glands; lateral veins 7-12-paired; petioles up to 1 cm long. Capitulescences terminal, corymbose. Capitula narrowly campanulate, 14-16 mm long, pedunculate. Receptacle convex, 1-1.5 mm in diam., hairy. Involucres slightly oblong-cylindrical, in 5-6 series, 8-10 mm long, 3-4 mm in diam. Phyllaries imbricate, green, margin piliferous, outer surface arachnoid without glands; the outer and the middle ones, ovate, apex obtuse; the inner ones lanceolate or ovate-lanceolate, apex obtuse or rounded. Florets 5-10; corollas funnelform, purple or white, glandular, corolla tubes 6-7 mm long; corolla lobes 4-4.5 mm long. Anthers 4-4.5 mm long, apical appendage acute, base obtuse. Styles purple, 9-10 mm long, branches 2, 4-5 mm long. Achenes turbinate, 3-3.5 mm long, 10-ribbed, covered with dense hairs and capitate glands. Pappus in 2 series of bristles, the inner ones 8.5-9 mm long, persistent.

Distribution.

Thailand: Chiang Mai, Chiang Rai. China (Yunnan), India, Nepal, Myanmar.

Specimens examined.

Thailand, Chiang Mai, Doi Chiangdao wildlife sanctuary, 19°24.11'N, 98°55.10'E, 20 Dec 2002, S. Bunwong 76 (KKU); Chiang Mai, Doi Ang Khang, 3 Jan 2008, 19°54.1'N, 99°2.4'E, 3 Jan 2008, S. Bunwong 378 (KKU, US); Fang, 11 Feb 1983, H. Koyama, H. Terao & Th. Wongprasert T-33345 (BKF); Fang, 12 Feb 1983, H. Koyama, H. Terao & Th. Wongprasert T-33458 (BKF); Mae Tang, Doi Chang, 12 Feb 1900, R. Pooma 382 (BKF); Doi Chang, 18 Jan 1983, H. Koyama, T. Yahara & W. Nanakorn T-32726 (BKF, L); Doi Chang, 23 Oct 1979, T. Shimizu, H. Toyokuni, H. Koyama, T. Yahama & T. Santisuk 20608 (BKF); Doi Anga Ga, Inthanon, 7 Feb 1931, H.B.G. Garrett 629 (BKF, BM, K, P); Doi Inthanon, 22 Mar 1967, T. Smitinand et al. 10280 (BK, BKF, K, L, P); Inthanon, 20 May 1970, Worawoot 95 (BKF); Doi Chiangdao, 7 Feb 1983, H. Koyama, H. Terao & Th. Wongprasert T-33185 (BKF); Chiang Rai, Doi Thung, Mae Sai, 14 Feb 1983, H. Koyama, H. Terao & Th. Wongprasert T-33515 (BKF).

Diagnostic characters.

This species is characterized by discinctly sweet smell, the habit of shrubs or subshrubs, and obtuse phyllaries.

Ecology.

Evergreen or pine-oak forest, alt. 1000-2000 m; flowering December to April.

Vernacular name.

Pim Pai Lin (พิมพ์ไพลิน).

Figures

Figure 9. Morphology of Vernonieae in Thailand 5. A-B Gymnanthemum extensum C Iodocephalopsis eberhardtii D Koyamasia calcarea E-F Koyamasia curtisii G Kurziella gymnoclada H Monosis parishii I Monosis volkameriifolia