Harmothoe goreensis Augener, 1918

Barnich, R. & Fiege, D., 2000, Revision of the Mediterranean species of Harmothoe Kinberg, 1856 Lagisca Malmgren, 1865 (Polychaeta: Polynoidae: Polynoinae) descriptions of a new genus and a new species, Journal of Natural History 34, pp. 1889-1938 : 1924-1925

Harmothoe goreensis Augener, 1918

(fi gures 1, 18A -D) Harmothoe goreensis Augener, 1918: 142 , pl. 2: fi gures 4-6, pl. 3: fi gure 42, text- fi gure 7; Day, 1967: 69, fi gure 1.9n -r.

Type material. Harmothoe goreensis , SYNTYPES: 2 spms., Westafrika , Ambrizette Angola), 12 m, leg. A. Hupfer (1889), det. Augener (1917), ZMH V 8651 (incomplete, no elytra) .

Additional material. 1 spm., Banyuls, Amphioxus -sand, det. as H. spinifera, ZMH 13997. 1 spm., September 1994, Banyuls, coralligeÁne, leg. D. Fiege, SMF 9235/1. spm., 12 October 1995, Banyuls, coralligeÁne, 24 m, leg. D. Fiege, SMF 9236/1. spms., 26 March 1997, Banyuls, coralligeÁne, 26 m, leg. D. Fiege and R. Barnich, SMF 9237/1. 7 spms., St. 129, 15 June 1950, Marseille, S de l’õÃle de Maire, 40 m, sable vaseux coÃtier, Coll. PeÂreÁs indeÂt. 1 spm., St. 188, 25 February 1951, Marseille, gravelle, det. as H. spinifera Ehlers, Coll. PeÂreÁs. 3 spms., St. 202, 22 May 1951, Marseille, fond de gravelle, det. as Lagisca extenuata (Grube) , Coll. PeÂreÁs. 1 spm.,. 212, 8 July 1951, Marseille, E de Planier, fonds vaseux coÃtiers, det as Harmothoe . spinifera Ehlers, Coll. PeÂreÁs. 5 spms., Gyf St. 306, 23 March 1953, Marseille, fond coralligeÁne du Cap Caveaux, Coll. PeÂreÁs indeÂt. 4 spms., 17 March 1955, Marseille, Riou, fond aÁ Peyssonnelia polymorpha , det. as Harmothoe imbricata L., Coll. PeÂreÁs. 6 spms., D1, 1976, Cortiou A.J., 86 m, det. as Harmothoe sp. by. Jojaruria, Coll. Bellan. 1 spm., F1, 1976, Cortiou A.J., 54 m, det. as Harmothoe . by A. Jojaruria, Coll. Bellan. 1 spm., Lesina (Italien), Adria , labelled Aphroditidae (Nychia) ’, Coll. Grube, ZMB F.1527.

Diagnosis. Anterior pair of eyes dorsolaterally on widest part of prostomium. Elytra: margin with short fringing papillae; surface covered by rounded conical

Revision of the Mediterranean Harmothoe and L agisca 1925 IG. 18. Harmothoe goreensis ((A) 17 March 1955, Marseille; (B -D) 9.94, Banyuls, SMF 9235/1). (A) Anterior end; dorsal view; (B) left middle elytron; (C) detail of posterior margin of same; (D) style of dorsal cirrus (scales 5 2 mm for A, B; 250 M m for C; 1 mm for D).

microtubercles. Dorsal and ventral tentacular cirri and dorsal cirri papillate, swelling subdistally then abruptly tapering to fi liform tip.

Description. Prostomium bilobed, with distinct cephalic peaks; median antenna with large ceratophore in anterior notch, style papillate, swelling subdistally then abruptly tapering to fi liform tip, pigmented basally and with two pigmented bands distally; lateral antennae with small ceratophores inserted ventrally and with short, papillate, tapering styles, each pigmented basally; palps papillate, longer than median antenna and tentacular cirri; anterior pair of eyes dorsolaterally on widest part of prostomium, posterior pair dorsally near hind margin. Tentaculophores inserted laterally to prostomium, each with two chaetae and a pair of papillate dorsal and ventral tentacular cirri, swelling subdistally then abruptly tapering to fi liform tip, each pigmented basally and with two pigmented bands distally. Second segment with fi rst pair of elytra, biramous parapodia, and long ventral buccal cirri, similar

tentacular cirri (fi gure 18A).

Fifteen pairs of elytra; elytral margin with short fringing papillae; surface covered

rounded conical microtubercles; pigmentation: mottled, brownish in alcohol

gures 1, 18B, C). Styles of dorsal cirri papillate, swelling subdistally then abruptly tapering to fi liform tip, extending beyond neurochaetae, each pigmented basally and with a pigmented band distally (fi gure 18D); styles of ventral cirri papillate, tapering, shorter than neuropodia.

Parapodia biramous, both rami with single aciculum penetrating epidermis; notopodia with short, rounded presetal and longer, pointed postsetal acicular lobe, neuropodia with subconical presetal acicular lobe with digitiform supra-acicular process and shorter rounded postsetal lobe. Notochaetae stouter than neurochaetae, 1926 with distinct rows of spines and blunt tips; neurochaetae with distinct rows of spines