Harpalus Latreille, 1802

Azadbakhsh, Saeed & Nozari, Jamasb, 2015, Checklist of the Iranian Ground Beetles (Coleoptera; Carabidae), Zootaxa 4024 (1), pp. 1-108 : 76-79

Genus Harpalus Latreille, 1802

Subgenus Semiophonus Schauberger, 1933

Harpalus (Semiophonus) signaticornis (Duftschmid, 1812)

Studied material: (ZMTU): Kolyak, 2008 (Prov. of Mazandaran) and Quamsar, 2012 (Prov. of Esfahan) Citations: Without exact locality ( Kataev & Wrase, 2003); Province East Azarbaijan ( Wrase, 2005).

Subgenus Cephalophonus Ganglbauer, 1891

Harpalus (Cephalophonus) cephalotes Fairmaire & Laboulbene, 1854 Citations : Aligudarz, Ghalikuh, 2300 m (Prov. of Lurestan; Wrase, 2009a).

Subgenus Pseudoophonus Motschulsky, 1844

Harpalus (Pseudoophonus) griseus (Panzer, 1796)

Studied material: Sirch. 2014 (Prov. of Kerman)

Citations: Kerman (Prov. of Kerman; Mandl, 1963); Khorramabad (Prov. of Mazandaran; Mandl, 1963); Tarsee, 2800 m (Prov. of Tehran; Mandl, 1963); Bardaksan, Mashhad, Dargaz, Serakhs, Ghaenat and Ferdows (Prov. of Razavi Khorasan; Sadeghi et al., 2010); Talesh (Prov. of Gilan; Salari et al., 2013).

Harpalus (Pseudoophonus) calceatus (Duftschmid, 1812)

Citations: Namin Village (Prov. of Ardabil; Atamehr, 2013).

Harpalus (Pseudoophonus) rufipes (Degeer, 1774)

Studied material: Khirood forest, 2012 (Prov. of Mazandaran), Mahan, 2014 (Prov. of Kerman).

Citations: Distributed in the North and center parts of Iran. Saguch (Prov. of Kerman; Mandl, 1963); Babol and Chalus (Prov. of Mazandaran; Mandl, 1963); Mashhad and Neyshabor (Prov. of Razavi Khorasan; Sadeghi et al., 2010); Mashahd C. (Prov. of Razavi Khorasan; Hosseini et al., 2012); Pars Abad Moghan Village (Prov. of Ardabil; Atamehr, 2013); Bandare Anzali, Fouman, Talesh and Roudbar (Prov. of Gilan; Salari et al., 2013).

Subgenus Cryptophonus Brandmayr et Zetto Brandmayr, 1982

Harpalus (Cryptophonus) litigiosus Dejean, 1829

Citations: Esfahan (Prov. of Esfahan; Kataev, 2013); Dezful (Prov. of Khuzestan; Kataev, 2013).

Harpalus (Cryptophonus) melancholicus melancholicus Dejean, 1829

Citations: Without exact locality (Hieke & Wrase, 1988); Without exact locality ( Kataev, 2013).

Harpalus (Cryptophonus) tenebrosus Dejean, 1829

Studied material: (ZMTU): Karaj, 2008 (Prov. of Elburz) and Kolyak, 2008 (Prov. of Mazandaran). Citations: Tehran (Prov. of Tehran; Kataev, 2013); Hasan Kiadeh (Prov. of Gilan; Kataev, 2013); 5 km North West of Ardakan, Sepidan (Prov. of Fars; Kataev, 2013); South East Khorasan and Kerman ( Kataev, 2013); Basmenj village (Prov. of East Azarbaijan; Atamehr, 2013).

Subgenus Harpalus Latreille, 1802

Harpalus (Harpalus) affinis (Schrank, 1781)

Studied material: Khirood forest, 2012 (Prov. of Mazandaran). Citations: Without exact locality (Hieke & Wrase, 1988).

Harpalus (Harpalus) amplicollis Ménétriés, 1848

Citations: Without exact locality ( Kataev & Wrase, 2003).

Harpalus (Harpalus) angulatus subangulatus Reitter, 1900

Citations: Gorgan env. (Prov. of Golestan; Afshari & Khormali, 2014).

Harpalus (Harpalus) antonowi (Tschitschérine, 1898)

Citations: Without exact locality ( Kataev & Wrase, 2003).

Harpalus (Harpalus) attenuatus Stephens, 1828

Citations: Without exact locality ( Kataev & Wrase, 2003).

Harpalus (Harpalus) caspius (Steven, 1806)

Studied material: (ZMTU): Kolijak, 2010 (Prov. of Mazandaran) and Lahijan, 2010 (Prov. of Gilan) Citations: Without exact locality ( Kataev & Wrase, 2003); Pars-Abad (Prov. of Ardabil; Samin et al., 2011).

Harpalus (Harpalus) convexus Faldermann, 1836

Studied material: (ZMTU): Damavand, 2600 m. 2008 (Prov. of Tehran), NowShahr, 2008 (Prov. of Mazandaran) and Bandar-e Anzali, 2008 (Prov. of Gilan).

Citations: Without exact locality ( Kataev & Wrase, 2003).

Harpalus (Harpalus) compressus Motschulsky, 1844

= amator Reitter, 1900

Citations: Mandl (1963) cited the species from Kuh-räng, 2500 m (Prov. of Esfahan) but this record is dubious as in the last Palearctic catalogue.

Harpalus (Harpalus) cupreus fastuosus Faldermann, 1836 Citations : Without exact locality ( Kataev & Wrase, 2003).

Harpalus (Harpalus) dimidiatus (P. Rossi, 1790)

Studied material: (ZMTU): Nowshahr and Kolijak (Prov. of Mazandaran).

Citations: Astrabad and Kopet Dag(Prov. of Golestan; Mlynar, 1979); Rudbarak and Kelardasht (Prov. of Mazandaran; Mlynar, 1979); Khorramabad (Prov. of Mazandaran; Mandl, 1963).

Harpalus (Harpalus) distinguendus distinguendus (Duftschmid, 1812)

Studied material: Kermanshah C., 2014. (prov, of Kermanshah); (ZMTU): Kordak River, 2008 (Prov. of Qazvin) and Hashtpar, 2012 (Prov. of Gilan).

Citations: Province Mazandaran ( Chaudoir, 1844); Mashhad, Torbat-e- Heydariye, Torbat-e- Jam, Kashmar and Ghaenat (Prov. of Razavi Khorasan; Sadeghi et al., 2010); Province Razavi Khorasan (Hosseini et al., 2012); Gorjan Village (Prov. of Ardabil; Atamehr, 2013).

Harpalus (Harpalus) famelicus famelicus Tschitschérine, 1898 Citations : Arak (Prov. of Markazi; Mandl, 1963).

Harpalus (Harpalus) flavicornis flavicornis Dejean, 1829

Citations: Without exact locality ( Kataev & Wrase, 2003); Ayuriq Village (Prov. of Ardabil; Atamehr, 2013).

Harpalus (Harpalus) fuscicornis Ménétriés, 1832

Studied material: (ZMTU): Gadaneh-e Heyran (Prov. of Ardabil).

Citations: Chalus env., Hasan Keif and Kelar Dasht, 600 m (Prov. of Mazandaran; Wrase, 2009a).

Harpalus (Harpalus) fuscipalpis Sturm, 1818 Studied material: Firooz Kola, 2010 (Prov. of Mazandaran). Citations: New record for Iran.

Harpalus (Harpalus) honestus (Dufischmid, 1812)

Citations: Without exact locality ( Kataev & Wrase, 2003); Chenaran (Prov. of Razavi Khorasan; Sadeghi et al., 2010).

Harpalus (Harpalus) hospes hospes Sturm, 1818

Citations: Samin et al. (2011) cited the species from Ahwaz (Prov. of Khuzestan), but this record is dubious because this species distributed in Northen countery of Iran and this record refer ro South of Iran.

Harpalus (Harpalus) kadleci Kataev & Wrase, 1995

Citations: Without exact locality ( Kataev & Wrase, 2003).

Harpalus (Harpalus) kazanensis Jedlička, 1958

Citations: Without exact locality ( Kataev & Wrase, 2003).

Harpalus (Harpalus) luteicornis (Dufischmid, 1812) Citations : Without exact locality ( Gueorguiev, 1995).

Harpalus (Harpalus) metallinus metallinus Ménétriés, 1836 Studied material: Kermanshah, 2014 (Prov. of Kermanshah) Citations: Sabzawaran (Prov. of Kerman; Mandl, 1963).

Harpalus (Harpalus) murzini Kataev, 2008

Citations: Rostamabad between Domash and Durfak, 1750–2000 m (Prov. of Gilan; Kataev, 2008).

Harpalus (Harpalus) oblitus oblitus Dejean, 1829

Citations: Kuh-räng (Prov. of Esfahan; Mandl, 1963); Without exact locality ( Kataev & Wrase, 2003); Mashhad, Sabzevar and Birjand (Prov. of Razavi Khorasan; Sadeghi et al., 2010).

Harpalus (Harpalus) reflexus anadoluensis Kataev, 1993 Citations : North west of Iran (Kataev, 1993).

Harpalus (Harpalus) rubripes (Dufischmid, 1812)

Studied material: Khirood Forest, 2012 (Prov. of Mazandaran); (ZMTU): Bandar-e Anzali, 2012 (Prov. of Gilan). Citations: Without exact locality ( Kataev & Wrase, 2003).

Harpalus (Harpalus) rufipalpis Sturm, 1818

Studied material: Kolijak, 1600-1850 m (Prov. of Mazandaran). Citations: New record for Iran.

Harpalus (Harpalus) saxicola Dejean, 1829

Citations: Without exact locality ( Kataev & Wrase, 2003).

Harpalus (Harpalus) serripes serripes (Quensel in Schonherr, 1806)

Studied material: (ZMTU): 15 km east of Baladeh, 2008 (Prov. of Mazandaran). Citations: Shahrivar Village (Prov. of Ardabil; Atamehr, 2013).

Harpalus (Harpalus) smaragdinus (Duftschmid, 1812)

Studied material: (ZMTU): Kolijak, 1600-1850 m (Prov. of Mazandaran). Citations: Without exact locality ( Kataev & Wrase, 2003); Ferdows (Prov. of Razavi Khorasan; Sadeghi et al., 2010).

Harpalus (Harpalus) smyrnensis medicus Kataev, 1993

Studied material: (ZMTU): Shemshak, 2008 (Prov. of Tehran), Kandovan, 2008 (Prov. of Mazandaran) and Harunie, 2012 (Prov. of Chahar Mahal va Bakhtiari).

Citations: Distributed mainly in north part of Iran. Without exact locality ( Kataev & Wrase, 2003); Gorgan env. (Prov. of Golestan; Afshari & Khormali, 2014).

Harpalus (Harpalus) smyrnensis raddei Tschitschérine, 1897 Citations : Kuh-räng, 2400 m (Prov. of Esfahan; Mandl, 1963).

Harpalus (Harpalus) subcylindricus Dejean, 1829

Citations: Without exact locality ( Kataev & Wrase, 2003).

Harpalus (Harpalus) tardus (Panzer, 1796)

Studied material: (ZMTU): Dohezar River and Javaher Deh, 2008 (Prov. of Mazandaran) and Astaneh, 2012 (Prov. of Gilan).

Citations: Without exact locality ( Kataev & Wrase, 2003).

Harpalus (Harpalus) zabroides Dejean, 1829

Studied material: (ZMTU): Damavand, Gosfand Sara, 2008 (Prov. of Tehran). Citations: Without exact locality ( Kataev & Wrase, 2003).

Harpalus (Harpalus) uniformis staudingeri Jedlièka, 1953

Citations: Quchan, 37°23´N, 58°35´E, 1921 m (Prov. of Razavi Khorasan; Kataev, 2015).