Helicopsyche (Petrotrichia) ambodiva, Johanson, Kjell Arne & Oláh, János, 2006

Johanson, Kjell Arne & Oláh, János, 2006, Eleven new species of Sericostomatoidea from Madagascar (Trichoptera: Helicopsychidae, Petrothrincidae, Sericostomatidae), Zootaxa 1205, pp. 1-30 : 4-6

Helicopsyche (Petrotrichia) ambodiva , new species

Figs 1–6 , 58

Helicopsyche . ambodiva is closely related to H. anomana Johanson, 2002 , especially in the general shape of the gonocoxite involving a short, inwardly oriented ventral branch. Helicopsyche ambodiva is distinguished by having the gonocoxite tapering more stepwise and not distally clavate; tergum X characteristically hooked; segment IX with the anterior margin triangular centrally (not dorsally as in anomana ), and the sternite of segment IX strongly produced posteriorly. In anomana segment IX is only slightly produced posteriorly.

Male head and thorax ( Figs 1, 2 ): Antennal scape about 4x as long as wide, slightly longer than diameter of compound eye. Maxillary palp ( Fig. 2 ) 2­segmented, 1st segment broadest, 2nd segment slightly longer than first. Interantennal warts separate from each other. Cephalic setal warts egg­shaped. Postantennal setal warts small, flat, rounded. Postocular setal warts long vertical. Pronotum with one pair of large, wide setal warts. Mesoscutum and mesoscutellum each with one pair of setal warts.

Male wings ( Fig. 3 ): Forewing length 3 mm, hind wing length 2,7 mm. Denuded wing membrane pale flavous with pale helvus veins. Venation similar to H. anomana Johanson and H. giboni Johanson. On forewing R5 originating at crossvein R3­R4. M and Cu separate longitudinal fold with sensillae. Discoidal cell long, about one fourth wing length. Abdominal reticulation absent. Sternal process on VI abdominal segment well developed.

Male abdomen and genitalia ( Figs 4–6 ): Segment IX well developed, anteriorly pointed triangularly, exactly in the middle ( Fig. 4 ); central part of posterior margin straight, sternite strongly produced posteriorly and almost rectangular ( Fig. 4 ); in ventral view ( Fig. 6 ) forming a separate ventral unit. Tergum IX very short, narrow in dorsal aspect ( Fig. 5 ). Superior appendage digitate, directed posterodorsally in lateral view ( Fig. 4 ) and slightly medially in dorsal view ( Fig. 5 ). Tergum X forming a pair of bifid elongate processes: mesially a short process, apically slightly hooked dorsally ( Fig. 4 ); laterally a much longer out­curving and tapering ventral branch slightly bending dorsally in latter half ( Fig. 4 ). Gonocoxite simple, a short ventral branch arising near base and generally oriented posterodorsally; main branch undulating, extending beyond ventral branch and narrowing in three steps. Basal plate not conspicuous. Phallic base broad due to the origin of the long dorsal process ( Fig. 4 ) running closely and almost parallel before crossing subapically ( Fig. 5 ); ventral process of phallus about 2x broader than dorsal process. Both dorsal and ventral processes slightly sinuous and down­curving along their lengths ( Fig. 4 ).

Etymology.The name is derived from the local name of the type locality of the species, Ambodivoangy.

Holotype male: MADAGASCAR: Maroantsetra, Ambodivoangy, iii. 1957 [R. Paulian] (MNHN, in alcohol).

Figures

FIGURE 58. Map of Madagascar showing distributions of Helicopsyche spp. (Helicopsychidae): H. ambodiva (circle); H. ninakosha and H. sahadika (rectangle); H. giboni (diamond); and H. anomana (triangle).

FIGURES 1 – 6. Helicopsyche (Petrotrichia) ambodiva, new species, holotype. 1 — head and thorax, dorsal; 2 — maxillary palp; 3 — right wing pair; 4 — genitalia, lateral; 5 — genitalia, dorsal; 6 — genitalia, ventral.