Helix (Helix) dormitoris KOBELT , 1898

Neubert, Eike, 2014, Revision of Helix LINNAEUS, 1758 in its eastern Mediterranean distribution area, and reassignment of Helix godetiana KOBELT, 1878 to Maltzanella HESSE, 1917 (Gastropoda, Pulmonata, Helicidae), Contributions to Natural History 26, pp. 1-200 : 133-136

Helix (Helix) dormitoris KOBELT, 1898 ( Figs 210–219 )

1898 Pomatia dormitoris KOBELT, Nachrichtsblatt der deutschen Malakozoologischen Gesellschaft 30: 163 [ Montenegro: Durmitor].

1898 Helix kolaschinensis KOBELT, Nachrichtsblatt der deutschen Malakozoologischen Gesellschaft 30: 164 [ Montenegro: Kolašin].

1906 Helix (Helicogena) kolasinensis ? var. bosnica KOBELT, Systematisches Conchylien-Cabinet VI : 263, Taf. 362, Fig. 9 , 10 [ Bosnia and Herzegovina: Bjelasnica bei Serajewo].

1906 Helix (Helicogena) stolacensis KOBELT, Systematisches Conchylien-Cabinet VI : 284, Taf. 374, Fig. 1–4 [Berg Stolac an der serbisch-bosnischen Grenze; =? 43° 47' 35N 19° 24' 22E, 1011 m alt.?].

1906 Helix (Helicogena) pivensis KOBELT, Systematisches Conchylien-Cabinet VI : 285, Taf. 374, Fig. 5–6 [ Montenegro: Durmitor gegen das Pivathal].

1906 Helix (Helicogena) pivensis lubnicensis KOBELT, Systematisches Conchylien-Cabinet VI : 286, Taf. 374, Fig. 7, 8 [ Bosnia and Herzegovina: Ljubicna].

1906 Helix (Helicogena) sturanyi KOBELT, Iconographie der Land- und Süsswassermollusken 2 (12): 42, Taf. 321, Fig. 2027 [ Bosnia and Herzegovina: Gacko].

1939 Helix dormitoris, Rasse arnautorum KNIPPER, Archiv für Naturgeschichte, N.F. 8: 351 [Aleksandrova vis (= Turcin) in der Šarplanina].

1939 Helix dormitoris, Rasse hajlensis KNIPPER, Archiv für Naturgeschichte, N.F. 8: 349 [Hajila Berg unweit Péc, Gipfelregion, 2000–2400 m; Serbia, Kosovo].

Type specimens:

dormitoris : lectotype SMF 9755 (SD Zilch, 1952: 152), H = 34.1, D = 32.75. kolaschinensis : lectotype SMF 9721 (SD Zilch, 1952: 153, err. under SMF 7921), H = 34.55, D = 33.0. bosnica: lectotype SMF 9757 (SD Zilch, 1952: 153), H = 34.7, D = 30.0. stolacensis : holotype SMF 9732, H = 47.2, D = 42.55. pivensis : holotype SMF 9727, H = 40.1, D = 37.05. lubnicensis: lectotype SMF 9788 (SD Zilch, 1952: 153), H = 34.8, D = 33.2. sturanyi : holotype SMF 9772, H = 35.8, D = 31.9. arnautorum: paratype SMF 60084, H = 33.3, D = 31.65. hajlensis: paratype SMF 60085, H = 37.4, D = 33.7.

Specimens examined:

Montenegro: Crno-Jezero , 1–2 Km from Zabljak, Durmitor, 43.1475 19.0876, 1440 m alt., 15.8.1987, NMBE 523963 / 8; Budecevica , bridge over the Tara, 43.1502 19. 2953, 690 m alt., 13.8.1987, NMBE 523962 / 189; Zabljak , Durmitor, 43.16 19.12, 27.5.2010, NMBE 23346 /3.

Other records:

Rudoka planina at the village Rudoka, 41.9344 20.8016; Neretva Valley close Jablanica (Hercegovina), 43.6602 17.7652; Rogatica, Bosnia, 43.7992 19.0048; Sarajevo, 43.8557 18.4088; village Pepice close Pec (not identified) (all Knipper 1939). — Cajnica, Bosnia, 43.5828 19.1012; Gorazda, Bosnia, 43.6691 18.9764; Boan-Tuschina, Montenegro (not identified) (all Hesse 1920: 221).

Diagnosis: shell medium sized, high conical spire, usually spiral No. 2 fused with No. 3, aperture cream to white.

Description: shell small to medium sized, thin, elongate spherical, high conical spire; basic shell colour greyish white with chestnut brown spirals, usually spiral No. 2 fused with No. 3 or all five spirals separate; protoconch large in comparison to shell, its diameter ranging from 5–6. 5 mm, smooth, white whorls; teleoconch of 3–4 whorls, smooth, with fine axial riblets and a few subsutural spiral incisions; last whorl only slightly descending towards the periphery; aperture rounded, apertural rim narrow, labial callus weak, cream to white coloured, columellar triangle small; umbilicus closed in adult shells.

Genital organs: There were no specimens available for dissection. Hesse (1920: 221, Taf. 658, Fig. 8, 9 . Taf. 659, Fig. 1 .) under H. kolaschinensis noted that the morphology of the genital organs is similar to those of H. pomatia , with the exception that all eight specimens investigated by him had a well developed diverticulum, which is small or even missing in H. pomatia .

Distribution ( Fig. 219 ): This species is a small range endemic to the Dinaric Alps in the Central Balkans.

Remarks: The status of this taxon as a separate species is not fully established, and the conchological differences to H. secernenda are few. In comparison to this species, H. dormitoris has smaller shells, with a more elevated spire, a relatively smaller and more rounded aperture, and an always white aperture. Knipper (1939: 349) records specimens of H. (Helicogena) kolasinensis ? var. bosnica from Sarajevo with a reddish columella, however, the type specimen SMF 9757 has a white aperture.

Figures

Figs 210–218. Helix (Helix) dormitoris KOBELT, 1898. Fig. 210 lectotype dormitoris SMF 9755, Durmitor, D = 32.75; Fig. 211 lectotype lubnicensis SMF 9788, Ljubicna, D = 33.2; Fig. 212 lectotype kolaschinensis SMF 9721, Kolašin, D = 33.0; Fig. 213 holotype pivensis SMF 9727, Durmitor gegen das Pivathal, D = 37.05; Fig. 214 holotype stolacensis SMF 9732, Berg Stolac, D = 42.55; Fig. 215 lectotype bosnica SMF 9757, Bjelasnica bei Serajewo, D = 30.0; Fig. 216 holotype sturanyi SMF 9772, Gacko, D = 31.09; Fig. 217 paratype hajlensis SMF 60085, Hajila Berg unweit Péc, D = 33.7; Fig. 218 paratype arnautorum SMF 60084, Aleksandrova vis, D = 31.65. — All photos Neubert, original size.

Fig. 219. Helix (Helix) dormitoris KOBELT, 1898. Distribution.

Figs 1–9. Helix (Helix) pomatia LINNAEUS, 1758. Fig. 1 compacta, syntype SMF 9573, D = 50.07, Hungary "Gestrüppe der Berglehnen"; Fig. 2 pulskyana, syntype SMF 9525/2a, Hungary "Inseln der Donau", D = 36.6; Fig. 3 hajnaldiana, syntype SMF 9658, Hungary "Budapest, im botanischen Garten", D = 34.1; Fig. 4 solitaria, syntype SMF 74708/2a, D = 31.1; Fig. 5 sabulosa, syntype SMF 9524, D = 40.5, Fig. 6 expansilabris, holotype SMF 9538, D = 45.5; Fig. 7 luteola, holotype SMF 9527, D = 34.6; Fig. 8 pannonica, lectotype SMF 9535, D = 40.5, Fig. 9 lednicensis, syntype SMF 9534/2a, D = 38.3. — All photos Neubert, original size.

Figs 10–19. Helix (Helix) pomatia LINNAEUS, 1758. Fig. 10 claudiensis, holotype SMF 9543, Klausenburg, D = 39.5; Fig. 11 dobrudschae, holotype SMF 9541, Rumania, Dobrudscha, Comana Vlasca, D = 46.2; Fig. 12 costellata, holotype SMF 9631, Markt in Bukarest, D = 39.7; Fig. 13 elsae, lectotype SMF 9529, nahe Bukarest, D = 34.9; Fig. 14 transsylvanica, holotype SMF 9634, Markt in Bukarest, D = 40.1; Fig. 15 christinae, lectotype SMF 9542, Runku über Bukarest, D = 44.5; Fig. 16 kapellae, holotype SMF 9544, Kapella Gebirge, D = 44.2; Fig. 17 serbica, holotype SMF 9540, Maidanek, D = 53.9; Fig. 18 thessalica, lectotype SMF 9568, Makrinitza bei Volos, D = 45.4; Fig. 19 rhodopensis, syntype SMF 9599/2a, Philippoppel, D = 41.0. — All photos Neubert, original size.