Heterotrigona itama ( Cockerell, 1918 )

Rasmussen, Claus, 2008, Catalog of the Indo-Malayan / Australasian stingless bees (Hymenoptera: Apidae: Meliponini), Zootaxa 1935, pp. 1-80 : 14-15

Heterotrigona itama ( Cockerell, 1918)

Trigona breviceps Cockerell 1919b: 244 : Holotype (BMNH 17b.1123, worker): examined, "Type" (red bor- der), "Sandakan / Borneo / Baker", " Trigona / breviceps / Ckll TYPE", "B.M. TYPE / HYM. / 17B.1123", "9591" (comparative notes, taxonomy); Type locality: MALAYSIA "Sandakan, Borneo (Baker, 9591)" (worker); Cockerell 1920b: 228 (distribution); MALAYSIA, Sandakan; Schwarz 1937: 292, 322-325*, 327, plate 3, 6 (key to species, taxonomy); Schwarz 1939a: 84, 96, 97, 98 (synonymy); Moure 1961: 203 (systematic position); Sakagami et al. 1990: 126 (synonymy).

Trigona itama Cockerell 1918: 387 : Holotype (USNM 29471, worker): examined (distribution, key to species); Type locality: SINGAPORE "Singapore" (unknown); Cockerell 1919b: 241-242, 244 (taxonomy, variation); MALAYSIA, Penang; Cockerell 1919c: 78, 79 (comparative notes, key to species); Cockerell 1919a: 121 (distribution, taxonomy, variation); INDONESIA, Pelaboean Ratoe, Java; SINGAPORE; Cockerell 1920a: 115 (comparative note); Cockerell 1925a: 492 (variation); MALAYSIA, Borneo, Sandakan; Cockerell 1927: 541 (distribution); MALAYSIA, Selangor, Pahang, near Genting, Sempat, Hill stream; Schwarz 1937: 283, 292, 319, 322-325*, 327, plate 3, 6 (key to species, taxonomy); Floral record: Melastoma ; Schwarz 1939a: 84, 85, 86, 90, 92, 95, 96-100*, 127, 133 (distribution, key to species, taxonomy); Floral record: Mimosa pudica ; Tephrosia candida ; Sandhouse 1943: 558 (citation); Schwarz 1948: 118, 119; Sakagami 1959: 119 (distribution); SINGAPORE; Moure 1961: 201, 203 (morphology, systematic position); Wille 1961: 117; Yoshikawa et al. 1969: 168, 172, 173, 174 (distribution, nest); MALAYSIA, Cameron Highland, Pahang; Fraser's Hill, Selagor-Pahang; George Town, Penang; Sabah, Jesselton; Sabah, Touaran; Sarawak, Kuching; Wille & Michener 1973: 14, 48 (nest); Sakagami 1975: 55 * (distribution, variation); MALAYSIA; THAILAND; Wille 1979b: 265 (comparative note); Appanah 1982: (pollination); MALAYSIA, Negri Sembiran, Pasoh Forest Reserve; Floral record: Xerospermum intermedium ; Sakagami et al. 1983b: 2-4, 13, 19, 20 (nest); Sakagami et al. 1985: (distribution, key to species); Inoue et al. 1985a: (aggressiveness, foraging, nest); Kiew 1985: (luring (unwashed shorts)); Inoue et al. 1985b: 45, 47, 48 (foraging); Baker & Delfinado-Baker 1985: 231 (inquiline ( Neocypholaelaps phooni : Ameroseiidae : Acari )); Atmowidjojo & Adisoemarto 1986: 227, 229 (pollination); Floral record: Indigofera ; Sakagami & Inoue 1987: 625 (comparative note); Uji 1987: (pollination); INDONE- SIA, E Kalimantan; Floral record: Nephelium lappaceum ; Camargo 1988: 370 (distribution, taxonomy); MALAYSIA, Kuala Lampur; SINGAPORE; Sakagami & Inoue 1989: 610, 613, 614, 616, 617 (morphology); Roubik 1989: 105, 133, 137, 156, 158; Michener 1990: 105, 120, 122, 125, 126, 127 (illustration, morphology, systematic position); Sakagami et al. 1990: (distribution, key to species); INDONESIA, Sumatra; Java; MALAYSIA, Borneo; THAILAND; Salmah et al. 1990: (distribution, nest (hollow trunks of clove ( Eugenia aromatica : Myrtaceae ); coconut; crevices)); INDONESIA, Sumatera Barat; Inoue et al. 1990b: (population); Inoue 1990: 582 (nest (tree cavities and others)); Michener & Roubik 1993: 260 (comparative note, morphology); Inoue et al. 1993: (nest); INDONESIA, Sumatra; Cunha 1994: (systematics); Tezuka & Maeta 1995: (foraging, pollen robbing); Roubik 1995: 151 (antropogenic nest); Camargo & Moure 1996: 109 (taxonomy (type series)); Salmah et al. 1996: (reproduction); Roubik 1996: 61, 62 (distribution); BRUNEI, Kuala Belalong; Michener 1997: 29 (type species); Nagamitsu & Inoue 1998: (foraging, morphology); Liow 1999: 106, 110, 127 (nest); Nagamitsu et al. 1999: (floral records (not by species), foraging); Heard 1999a: 196 (citation); Michener 2000: 801- 802; Chinajariyawong & Saiboon 2000: (distribution, nest); THAILAND, Songkhla; Amano et al. 2000: (citation); Liow et al. 2001: 184; Costa et al. 2003: (systematics); Eltz 2004: 322 (abundance);

Samejima et al. 2004: (population); MALAYSIA, Sarawak, Miri, Upper Baram area; Nagamitsu & Inoue 2005: 76, 83, 84 (distribution, foraging); Chinh et al. 2005: 368 (citation); Klakasikorn et al. 2005: 6 (citation); Roubik 2006: 131 (nest); Rasmussen & Cameron 2007: (systematics); Leonhardt et al. 2007: (foraging); Floral record: Averrhoa carambola ; Cassia fistula ; Michener 2007a: 826-827; Wattanachaiyingcharoen & Jongjitvimol 2007: 73 (comparative note).

Heterotrigona indet. Roubik 2006: 127, 132 (nest).