Incasarcus ochoai Kury & Maury, 1998

Benedetti, Alipio Rezende & Pinto-da-Rocha, Ricardo, 2022, Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species, Arthropod Systematics & amp; Phylogeny 80, pp. 309-388 : 309

3.33. Incasarcus ochoai Kury & Maury, 1998

Figs 4D , 9J-N , 18D-F , 30

Incasarcus ochoai Kury & Maury, 1998: 152 (desc.), 160 (key), figs. 21 (male dorsal habitus, chelicerae, pedipalpus, trochanter-patella IV), 22 (penis dorsal view); 23 (penis lateral view), 24 (male lateral habitus, chelicera, pedipalpus), 25 (female dorsal habitus), 26 (male sternum, coxae I-IV); 27 (male trochanter-patella IV), 28-31 (tarsi I-IV); Kury 2003: 144 (cat.); Ázara et al. 2020: 476 (cit), fig. 2 (cladogram).

Redescription.

MALE: Measurements (n =3) DSW: 5.1-5.2 (5.2); DSL: 6.3-6.5 (6.5); CL: 1.6-1.7 (2.6). FIVL: 11.0-11.2 (11.2). ChL: 3.5-4.7 (3.5). Coloration (in ethanol): Yellow with black spots covering the areas of the dorsal scutum; pedipalpus and chelicerae brown. Legs I-IV yellow. Dorsum: (Fig. 4D ) Median elevation of anterior margin of dorsal scutum with granules. Ocularium with well-marked median depression; sparse granules. Carapace with sparse granules. Areas I-IV with granules more densely distributed than on carapace; unarmed. Lateral margins of dorsal scutum with granules throughout their length. Posterior margin of dorsal scutum slightly granulate. Free tergites I-III with a row of granules. Chelicerae: (Fig. 4D ) Swollen in large males, slightly greater than those on females in smaller males (as in the holotype). Segment I granulate. Segment II with small granules; finger with two teeth. Segment III with two teeth. Pedipalpus : With granules scattered throughout the dorsal portion of the femur-patella. Trochanter with a ventroapical setiferous tubercle. Femur with a ventrobasal setiferous tubercle; a ventral row of seven setiferous tubercle throughout all extension of femur, except the apex and base; one proapical spine. Patella with a small proapical tubercle. Venter: Coxae I-IV with granules; Coxae I-II with 2-3 sparse tubercles. Rows of tubercles between the coxae II-III and III-IV. Genital area and free sternites few granulate. Anal operculum sparsely granulate. Legs: (Figs 4D , 9J-N ) Coxa I-II each with a retrolateral and a prolateral apophysis. Coxa III unarmed. Coxa IV with granules sparsely scattered. Trochanters I-IV unarmed and granular. Femora I-III with sparse granules. Femur IV with small granules throughout its surface; one retroventral row of 23-24 acuminate tubercles, the basal larger (in large males, the tubercles are larger and more robust); a prodorsal row of 7-8 tubercles at the base region (in large males, the tubercles are large and acuminated, whereas in smaller males, this row is virtually inconspicuous); a prolateral row of 5-7 tubercles, present only in the middle portion of femur, with some tubercles closer to the apex (only in large males). Patellae I-III granulate. Patella IV with a ventral row of five acuminate tubercles along the basal ½ (in smaller males, tubercles are smaller). Tibiae I-IV granulate. Tarsal segmentation: (n =3) 8-10 (10), 18-19 (19), 9-11 (10), 10-13 (13). Penis: (Fig. 18D-F ) VP subrectangular, distal margin slightly convex; with conspicuous lateral projections; lateral flanges ventrally folded. MS C1-C5 subapical long and curved; MS A1 median long and straight (half length of MS C), and MS A2-A3 sub basal long and straight; MS B1 basal short (placed near lateral sacs); MS D1 short, placed on flange near to MS C; MS E1-E2 short (larger than MS B and MS D), placed on ventrally fold of flanges. Lateral sacs long, apically slightly acuminated, with long T3-like microsetae. Stylus with broad apex, with a long ventral projection with small projections throughout its extension. Promontory straight. - FEMALE: Measurements (n =3) DSW: 5.2-5.7; DSL: 5.9-6.2; CL: 1.5. FIVL: 10.1-10.7. ChL: 1.2--1.3. Chelicerae not swollen. Femur IV unarmed. Tarsal segmentation (n =3): 8-9, 13-16, 9-10, 10-12.

Diagnosis.

It differs from other species of the genus by the set of following characters: DS without silver-white coloration; DS granulate; ocularium, areas I-IV and free tergites I-III unarmed (Fig. 4D ); male femur IV with one ventral row of 23-24 acuminate tubercles, a prolateral row of 5-7 tubercles and a prodorsal row of 7-8 tubercles (virtually inconspicuous in smaller males; Fig. 9J-N ).

Distribution.

(Fig. 30 ) PERU. Cusco. Urubamba province.

Material examined.

Type material: Holotype ♂, ' PERU, Cusco, Urubamba Province, Huayllabamba district, Yanacocha , Huayocari , Huayoccare , 3,000-4,000 m a.s.l., 13°20′S 72°02′W, 14/XI/1992, J.C. Chaparro leg. (MACN 9551) - Paratype ♀, ‘ditto’ (MACN 9552); Paratypes 2 ♀, ' PERU, Cusco, Urubamba Province, Huayllabamba district, Yanacocha , Huayocari , Huayoccare , 3,000-4,000 m a.s.l., 13°20′S 72°02′W | 17/VI/1995, J.A. Ochoa leg. (MACN 9553) ; Paratype 2 ♂, ‘ditto’ (MACN 9554); Paratype 1 ♂, ‘ditto’ (MNRJ 5401); Paratype 1 ♂, ‘ditto’ (MNRJ 5402 ♀).

Figures

Figure 4. Habitus, dorsal of males of Huancabamba gen. nov. and Incasarcus. A H. kubricki gen. et sp. nov.; B I. argenteus Kury & Maury, 1998; C I. dianae Kury & Maury, 1998; D I. ochoai Kury & Maury, 1998; E I. pictus Kury & Maury, 1998; F I. viracocha Kury & Maury, 1998; Legend bars = 1 mm.

Figure 9. Male trochanter-patella / tibia IV of Incasarcus. A - B Trochanter-tibia IV of I. argenteus Kury & Maury, 1998, dorsal and ventral view, respectively; C - D Trochanter-tibia IV of I. dianae Kury & Maury, 1998, dorsal and ventral view, respectively; E-F Trochanter-tibia IV of I. pictus Kury & Maury, 1998, dorsal and ventral view, respectively; G - H Trochanter-tibia IV of I. viracocha Kury & Maury, 1998, dorsal and ventral view, respectively; I Trochanter-patella of I. viracocha, lateral view; J Trochanter-patella of I. ochoai Kury & Maury, 1998, large male, dorsal view; K - L Trochanter-tibia of I. ochoai, small male, dorsal and ventral view, respectively; M Trochanter-patella of I. ochoai, large male, lateral view; N Trochanter-patella of I. ochoai, small male, lateral view; Legend bars = 1 mm.

Figure 18. Penis of Huancabamba gen. nov. and Incasarcus, dorsal, lateral and ventral views, respectively. A - C H. kubricki gen. et sp. nov.; D - F I. ochoai Kury & Maury, 1998; G - I I. viracocha Kury & Maury, 1998; Legend bars = 0.1 mm.

Figure 30. Distribution of Incasarcus in Cusco, Peru.