Kolbasinella Khramov, 2014a

Winterton, Shaun L., Martins, Caleb Califre, Makarkin, Vladimir, Ardila-Camacho, Adrian & Wang, Yongjie, 2019, Lance lacewings of the world (Neuroptera: Archeosmylidae, Osmylidae, Saucrosmylidae): review of living and fossil genera, Zootaxa 4581 (1), pp. 1-99 : 22

Kolbasinella Khramov, 2014a

Type species. Kolbasinella elongata Khramov, 2014a: 304 (by original designation).

Diagnosis. Hind wing narrow; RP1 originating at approximately 1/5 wing length; RP with 12 branches MA pectinately branched near wing margin (7 closely spaced branches); numerous regularly arranged crossveins throughout wing, outer gradate series poorly formed near wing margin; MP with only a few pectinate branches; CuA strongly pectinate (ca. 19 very short branches); FW unknown.

Comments. Kolbasinella was placed in Gumillinae by Khramov (2014a) due to the crossvein density and lack of trichosors in the wing margin. While these are indeed traits exhibited (in part) by Gumillinae , they are not diagnostic of the subfamily; some Gumillinae do have trichosors in the distal part of the wing (e.g., Gumilla ). Unfortunately, since Kolbasinella is only known from a hind wing, the justification presented by Khramov (2014a) for placement in Gumillinae is equivocal as important diagnostic antennal and forewing features are absent. The hind wing of Kolbasinella does not show the vein CuP but based on the shape of the other wing veins and the wing margin, it suggests that CuP is at least not elongate and pectinately branched, therefore placing it in one of three subfamilies, Spilosmylinae , Protosmylinae or Gumillinae . However, the genus is tentatively retained in Gumillinae here as its hind wing does not possess the basal sinuate crossvein 1r-m, a state diagnostic for the subfamily (1r-m is present in Spilosmylinae and Protosmylinae ). The lack of end-twigging in the posterior part of the wing is typical of Gumillinae but could also be due to the elongate shape of the wing and the proximity of the longitudinal veins (i.e., M and Cu) to the wing margin.