Landouria naggsi, Nurinsiyah & Neiber & Hausdorf, 2019

Nurinsiyah, Ayu Savitri, Neiber, Marco T. & Hausdorf, Bernhard, 2019, Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java, European Journal of Taxonomy 526, pp. 1-73 : 27-29

Landouria naggsi sp. nov.

urn:lsid:zoobank.org:act:4EF9E468-DDEF-460D-939C-7D9C86254CF4

Figs 25 , 52 ; Tables 3–4

Diagnosis

Landouria naggsi sp. nov. is characterized by an angular, tightly coiled, high conical shell with scales all over the teleoconch.

Etymology

Landouria naggsi sp. nov. is named in honour of the British malacologist Fred Naggs, who brought the authors of this study together.

Material examined

Holotype

INDONESIA • holotype; West Java, Cibitung, near Cikaso Waterfall , teak agroforestry; 7°21′41″ S, 106°37′10″ E; 47 m a.s.l.; 5 Jul. 2015; A.S. Nurinsiyah and T. Setiadi leg.; D = 11.8 mm, H = 9.8 mm; MZB 20424 .

Paratypes

INDONESIA – West Java • 1 spec.; Cibitung, near Cikaso Waterfall , riparian (high shrubs); 7°21′40″ S, 106°37′05″ E; 61 m a.s.l.; MZB 20426 • 1 spec.; same data as for preceding; ZMH 148184 • 3 spec.; same data as for holotype; MZB 20425 • 2 spec.; same data as for holotype; ZMH 148183 .

Description

SHELL (Fig. 25; Tables 3–4 ). Conical, with 6.25–7 slightly convex whorls; protoconch almost smooth; teleoconch with irregular wrinkles; incised spiral lines not discernible; scales all over teleoconch; tuberculate around umbilicus; brownish-corneous; body whorl angular; aperture oval; upper insertion of peristome slightly descending; peristome expanded, reflexed and slightly thickened; umbilicus hardly eccentric, wide, comprising 28–33% of shell diameter, hardly obscured by columellar edge.

GENITALIA. Unknown.

Remarks

Landouria naggsi sp. nov. resembles L. parahyangensis sp. nov. (Fig. 48), from which it differs in the smaller (D 11.6–14.0 mm vs 14.7–16.9 mm in L. parahyangensis sp. nov.), higher conical (D/H: 1.21– 1.37 vs 1.69–1.94 in L. parahyangensis sp. nov.), less sharply keeled shell with a narrower umbilicus (U/D 0.28–0.33 vs 0.39–0.45 in L. parahyangensis sp. nov.) and in having scales all over the teleoconch.

With regard to the conical shell form L. naggsi sp. nov. resembles L. conoidea ( Leschke, 1914) ( Fig. 4 ). For differences, see remarks under that species.

Distribution

Landouria naggsi sp. nov. is known only from the surroundings of Cikaso Waterfall in Sukabumi Regency (Fig. 52).

Figures

Figs 25–28 (opposite page). Shells of species of Landouria from Java. 25. L. naggsi sp. nov., West Java, Cibitung, near Cikaso Waterfall (holotype, MZB 20424). 26. L. epiplatia (Möllendorff, 1897), West Java, Jampang (lectotype of Plectotropis epiplatia, SMF 8909). 27. L. epiplatia, West Java, Jampang (lectotype of Plectotropis trichotrochium Möllendorff, 1897, SMF 8918). 28. L. epiplatia?, West Java, Bogor, Gunung Gede (ZMH 79946). Scale bar: 5 mm.

Figs 52–55 (opposite page). Distribution of species of Landouria in Java (10′ grid). 52. L. winteriana (Pfeiffer, 1842), L. naggsi sp. nov., L. nusakambangensis sp. nov., L. petrukensis sp. nov., L. tholiformis sp. nov., L. tonywhitteni sp. nov., L. smimensis (Mousson, 1848). 53. L. conoidea (Leschke, 1914), L. abdidalem sp. nov., L. sewuensis sp. nov., L. sukoliloensis sp. nov., L. pacitanensis sp. nov., L. intumescens (Martens, 1867), L. madurensis sp. nov. 54. L. rotatoria (Pfeiffer, 1842), L. nodifera sp. nov., L. parahyangensis sp. nov., L. schepmani (Möllendorff, 1897), L. zonifera sp. nov., L. ciliocincta (Möllendorff, 1897), L. pakidulan sp. nov. 55. L. epiplatia (Möllendorff, 1897), L. monticola van Benthem Jutting, 1950, L. davini Dharma, 2015, L. menorehensis sp. nov., L. dharmai sp. nov., L. moussoniana (Martens, 1867).

Figs 2–5. Shells of species of Landouria from Java. 2. L. winteriana (Pfeiffer, 1842), West Java, Padalarang, Gunung Masigit (neotype of Helix winteriana, ZMA 376193a). 3. L. winteriana, West Java, Cibodas (syntype of Plectotropis kraepelini Leschke, 1914, ZMH 98416). 4. L. conoidea (Leschke, 1914), West Java, Bogor (syntype, ZMH 98144). 5. L. rotatoria (Pfeiffer, 1842), West Java, Gunung Salak (neotype of Helix rotatoria, MZB 19205). Scale bar: 5 mm.