Lecanorchis ohwii Masamune (1933: 208)

Suetsugu, Kenji, Hsu, Tian-Chuan & Fukunaga, Hirokazu, 2017, Lectotypification of Lecanorchis ohwii (Vanilleae, Vanilloideae, Orchidaceae) with discussions of its taxonomic identity, Phytotaxa 309 (3), pp. 259-264 : 260-263

Lecanorchis ohwii Masamune (1933: 208) .

Type: — TAIWAN. New Taipei City: Wulai District, “inter Tyakon et Rarasan” (between Zhakong and Mt. Lala), 13 April 1933, J. Ohwi

847 (lectotype designated here, KYO-00022233). Synonyms: — Lecanorchis cerina var. albida Lin (1987: 147 , as “ albidus ”) ≡ Lecanorchis cerina f. albida (T.P. Lin) Ying (1990: 562)

≡ Lecanorchis albida (T.P. Lin) Lin (2016: 103) ≡ Lecanorchis kiusiana var. albida (T.P. Lin) Hsu (2016: 100) , syn. nov. Type: —

TAIWAN. New Taipei City, Mt. Pataoerh. 1100 m, April 1, 1979, T. P. Lin 439 (holotype TAIF 74168), T. P. Lin 414 (paratype TAIF

127050). Lecanorchis cerina auct . non Fukuy.: Lin (1987: 144), pro parte. Lecanorchis japonica auct . non Blume (1856: 188): Su (2000: 934), pro parte. Lecanorchis kiusiana auct . non Tuyama (1955: 182): Chung (2008: 50). Lecanorchis nigricans auct . non Honda: Liu & Su (1978: 1039), pro parte.

Distribution: — Taiwan.

Additional specimens examined: — TAIWAN. Hualien County: Tatung , 800–900 m, 9 April 2009, Chung 9537 ( TAIF), 800–900 m, 16 April 2011, Hsu 3874 ( TAIF). Nantou County, Shuishetashan, 1200–1300 m, 11 April 2013, Shen s.n. ( TAI). New Taipei City: Chiuchiungken, 300–400 m, 25 April 2011, Hsu 3938 ( TAIF) ; Mt. Pataoerh ( Pataoergshan ), 900–1000 m, 31 March 2007, Hsu 736 ( TAIF) ; 600–800 m, 11 April 2015, Hsu 7548 ( TAIF) ; Mt. Tapaoke , 700–800 m, 14 April 2007, Hsu 762 ( TAIF) ; Togokei , 800 m, 1 May 1936, Suzuki s.n. ( KPM) ; 900 m, 3 May 1936, Suzuki s.n. ( KPM). Pingtung County: Mt. Lilung ( Lilungshan ), 800–900 m, 26 February 2005, Hsu 303 ( TAIF) ; ca. 800 m, 9 March 1999, Wu 1054 ( HAST, TNM) .

Taxonomic remarks: — Lecanorchis ohwii can be diagnosed by the following combination of characters: inflorescences always simple aboveground, 10–30 cm tall; scape ± soft and fleshy in flowering; sepals 14–19 mm long, semi-spreading; lip 13–18 mm long, obviously 3-lobed; side-lobes of lip acute at apices, entire in margins; mid-lobe of lip elliptic-rectangular, ± recurved, 5–7 mm long and wide when flattened; hairs on mid-lobe of lip densely papillose near apices ( Fig. 2Q ); column 10–12 mm long, with well-developed column wings nearly entirely fusing with clinandrium ( Fig. 2J–L ). Correct descriptions and illustrations of this species could also be seen in Lin (1987: 147–149; pl. 85–88, as L. cerina var. albidus ), Chung (2008: 50, as L. kiusiana ; 2015: 255, as L. cerina var. albidus ) and Hsu & Chung (2016: 100, as L. kiusiana var. albida ).

A key to L. ohwii and taxonomic confusing or morphologically similar taxa in E Asia is also provided as follow:

1. Inflorescences sometimes branched aboveground; lip unlobed, obviously cucullate ( China, Japan, and Taiwan) ......... L. nigricans

- Inflorescences always simple aboveground; lip obvioously 3-lobed, not cucullate..........................................................................2

2. Inflorescences 30–60 cm high; sepals 25–30 mm long; lip 20–23 mm long; column 14–15 mm long (Taiwan) ................ L. cerina

- Inflorescences 10–45 cm tall; sepals 14–20 mm long; lip 13–18 mm long; column 10–12 mm long...............................................3

3. Lip not papillose along margins; multicellular hairs on mid-lobe of lip with glabrous surfaces ( Japan) ......................... L. japonica

- Lip minutely papillose along margins; multicellular hairs on mid-lobe of lip densely papillose near apices...................................4

4. Sepals hardly spreading; side-lobes of lip rounded at apex and denticulate in margin; mid-lobe of lip ca. 4 mm long; column distinctly trifid at apex, with column wings partially free from clinandrium ( Japan and Korea) ...................................... L. kiusiana

- Sepals semi-spreading; side-lobes of lip acute at apex and entire in margin; mid-lobe of lip 5–7 mm long; column subtruncate or obscurely trifid at apex, with column wings nearly entirely fusing with clinandrium (Taiwan) ............................................ L. ohwii

Figures

FIGURE 2. Morphology of Lecanorchis ohwii (based on Hsu 736). A. Habitat and habits. B & C. Flower (B, front view; C, lateral view). D. Dorsal sepal. E. Petal. F. Lateral sepal. G & H. Lip and column (G, dorsal view; H, lateral view). I. Lip. J–L. Column (J, lateral view; K, ventral view; L, oblique ventral view). M–P.Anther (M, dorsal view; N, lateral view; O, ventral view; P, oblique ventral view). Q. Hairs adaxially on midlobe of the lip, showing the densely papillate apices. Scale bars: B–L = 1 cm; M–Q = 1 mm. Photographed by T.C.Hsu.