Leptogamasus (Valigamasus), Karg, 1993

Bolger, Thomas, Arroyo, Julio & Piotrowska, Katarzyna, 2018, A catalogue of the species of Mesostigmata (Arachnida, Acari, Parasitiformes) recorded from Ireland including information on their geographical distribution and habitats, Zootaxa 4519 (1), pp. 1-220 : 53-90

Subgenus Valigamasus Karg, 1993

• Leptogamasus (Valigamasus) pannonicus (Willmann, 1951) • Original name: Pergamasus pannonicus Willmann, 1951 . • In: Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch -Naturwissenschaftliche

Classe. Abt, 160: 97.

Irish Records: Organic horizon under Sitka spruce ( Arroyo et al., 2008).

Counties: Laois.

Lysigamasus Karg, 1971

• Lysigamasus armatus Halbert, 1915

• Original name: Gamasus runciger var. armatus Halbert, 1915 .

• In: Proceedings of the Royal Irish Academy, 31: 51.

• Combinations/variations used in Irish literature: Pergamasus armatus Halbert : Cunningham, 1975; Ganley,

1976; Curry, 1976a; Purvis, 1978; Breslin, 1979; Curry, 1979; O’Neill, 1981; Heneghan, 1994; O’Connell,

1994), Paragamasus armatus Halbert ( Curry & O’Neill, 1979; Bolger, 1980; Purvis, 1982; Skorupski & Lux-

ton, 1998), Pergamasus (Paragamasus) armatus Halbert ( Whelan, 1976) .

Irish Records: Under bark of decaying fir tree and in rotten wood ( Halbert, 1915); pasture ( Cunningham, 1975; Curry, 1976a; Ganley, 1976; Purvis, 1978; Curry & O’Neill, 1979; Bolger, 1980; O’Neill, 1981; Piotrowska, 2009); pasture on peat ( Whelan, 1976); manure on soil surface (Curry, 1979); soil of sugar beet crop ( Breslin, 1979); soil under birch, horse chestnut, oak and pine (unpublished data); Iris bog at seashore ( Purvis, 1982); soil under Norway spruce ( Heneghan, 1994); soil in broadleaved woodland, Leyland cypress, Western hemlock, Monterey pine and pasture (unpublished data); woodland, rough grassland, under bracken, Sphagnum , cliff (Bolger, in press); fungal sporocarp ( O’Connell, 1994, unpublished data); moss on Sitka spruce ( Arroyo et al., 2010).

Counties: Carlow, Cork, Dublin, Galway, Kildare, Kilkenny, Laois, Mayo, Meath, Wexford, Wicklow.

• Lysigamasus celticus ( Bhattacharyya, 1963)

• Original name: Pergamasus (Paragamasus) celticus Bhattacharyya, 1963 .

• In: Bulletin of the British Museum (Natural History) Zoology, 11: 192.

• Combinations/variations used in Irish literature: Pergamasus celticus Bhattacharyya ( Curry, 1969a; Ganley,

1976; Heneghan, 1994; O’Connell, 1994; Heneghan & Bolger, 1996a), Paragamasus celticus (Bhattacharyya)

( Purvis, 1982; Skorupski & Luxton, 1998).

Irish Records: Pines and rhododendrons ( Bhattacharyya, 1963); pasture ( Curry, 1969a; Ganley, 1976; unpublished data); Iris bog at seashore ( Purvis, 1982); soil under Norway spruce ( Heneghan, 1994; Heneghan & Bolger, 1996a, unpublished data); pasture, rough grazing, coniferous forest and arable soils ( Arroyo et al., 2009); nest of Lasius flavus and pasture ( Arroyo et al., 2015); arable soil, wet pasture; rough grassland, cliff, woodland (Bolger, in press); fungal fruiting body (unpublished data); fungal sporocarp ( O’Connell, 1994); moss on soil under oak, ash and Scots pine; soil under Sitka spruce, Scots pine ( Arroyo et al., 2012, 2013a); fixed and fore dunes of Machair system ( Arroyo & Bolger, 2011); riverbank (unpublished data); Atlantic blanket bog, raised bog and fen ( Wisdom, 2010; Wisdom et al., 2011).

Counties: Clare, Cork, Dublin, Galway, Kerry, Kildare, Kilkenny, Laois, Limerick, Mayo, Offaly, Roscommon, Sligo, Tipperary, Wexford, Wicklow.

• Lysigamasus conus (Karg, 1971)

• Original name: Pergamasus conus Karg, 1971 .

• In: Die Tierwelt Deutschlands, 59: 378.

Irish Records: Soil under birch, oak and mixed broadleaves (unpublished data).

Counties: Dublin, Wexford.

• Lysigamasus digitulus (Karg, 1963)

• Original name: Pergamasus digitulus Karg, 1963 .

• In: Deutsche Entomologische Zeitschrift, 10: 78.

• Combinations/variations used in Irish literature: Pergamasus cambriensis Bhattacharyya ( Curry, 1969a;

Ganley, 1976; Emmanuel, 1977; Whelan 1986; O’Connell, 1994, Little & Bolger, 1995), Pergamasus

(Paragamasus) cambriensis Bhattacharyya ( Whelan, 1976, 1978, 1985), Paragamasus digitulus Karg ( Purvis,

1982; Skorupski & Luxton, 1998).

Irish Records: Pasture ( Curry, 1969a; Ganley, 1976; Piotrowska, 2009); pasture on peat ( Whelan, 1976, 1978, 1985, 1986); hedgerow around and soil in barley field ( Emmanuel, 1977); pasture, gorse bank and Salix ditch at seashore ( Purvis, 1982); ash and spruce woodland and pasture (unpublished data, Little & Bolger, 1995); fungal sporocarp ( O’Connell, 1994); riverbank (unpublished data).

Counties: Cork, Dublin, Kildare, Wexford.

• Lysigamasus femoratus ( Bhattacharyya, 1963)

• Original name: Pergamasus (Paragamasus) femoratus Bhattacharyya, 1963 .

• In: Bulletin of the British Museum (Natural History) Zoology, 11:197.

• Combinations/variations used in Irish literature: Pergamasus femoratus Bhattacharyya ( Curry, 1969a, b,

1973; Cunningham, 1975; Purvis, 1978; Tuohy, 1979; Bolger, 1985; McCarthy, 2004), Paragamasus femoratus

Bhattacharyya (Bolger, 1980).

Irish Records: Moss on lakeside rocks ( Bhattacharyya, 1963); pasture ( Curry, 1969a, b, 1973; Cunningham, 1975; Purvis, 1978; Tuohy, 1979; Bolger, 1980, unpublished data); Salix litter on peat ( Bolger, 1985); in soil under Norway spruce ( McCarthy, 2004).

Counties: Clare, Kildare, Kilkenny, Meath.

• Lysigamasus lapponicus (Trägårdh, 1910)

• Original name: Pergamasus lapponicus Trägårdh, 1910 .

• In: Naturwissenschaften Untersuchungen Sarekgeberg, 4: 408.

• Combinations/variations used in Irish literature: Gamasus (Pergamasus) lapponicus Trägårdh ( Halbert, 1915; Hull, 1918, Valarasan, 1971, Pergamasus lapponicus (Trägårdh) ( Ganley, 1976; O’Connell, 1994; O’Connell & Bolger, 1997b), Pergamasus (Paragamasus) lapponicus Trägårdh ( Whelan, 1976) , Pergamasus lapponicus Trägårdh ( Curry, 1969a, b, 1973; Purvis, 1978) Paragamasus lapponicus (Trägårdh) ( Breslin, 1979; Purvis, 1982; Skorupski & Luxton, 1998).

Irish Records: Under bark and in fir cones ( Halbert, 1915); lakeside ( Halbert, 1915); pasture ( Curry,

1969a, b, 1973; Valarasan, 1971; Ganley, 1976; Purvis, 1978; Piotrowska, 2009); pasture on peat ( Whelan,

1976); soil of sugar beet crop ( Breslin, 1979); Iris bog, bracken bed and Salix ditch at seashore ( Purvis,

1982); rough grassland, cliff (Bolger, in press); fungal sporocarp ( O’Connell, 1994; O’Connell & Bolger,

1997b, unpublished data); soil under Norway spruce ( Heneghan, 1994, unpublished data); soil in spruce

and broadleaved woodland (unpublished data).

Counties: Carlow, Dublin, Galway, Kildare, Kilkenny, Mayo, Wexford, Wicklow.

• Lysigamasus londonensis ( Bhattacharyya, 1963) • Original name: Pergamasus (Paragamasus) londonensis Bhattacharyya, 1963 . • In: Bulletin of the British Museum (Natural History) Zoology, 11: 207. • Combinations/variations used in Irish literature: Pergamasus londonensis Bhattacharyya ( Ganley, 1976) .

Irish Records: Pasture ( Ganley, 1976); soil under Sitka spuce and mixed broadleaved and coniferous

stands (unpublished data).

Counties: Dublin, Wexford.

• Lysigamasus misellus (Berlese, 1904)

• Original name: Gamasus (Pergamasus) misellus Berlese, 1904 .

• In: Redia 1: 237.

• Combinations/variations used in Irish literature: Pergamasus misellus Berlese ( Curry, 1969a; Ganley, 1976;

Breslin, 1979), Paragamasus misellus (Berlese) (Bolger, 1980; Purvis, 1982; Skorupski & Luxton, 1998).

Irish Records: Pasture ( Curry, 1969a; Ganley, 1976; Bolger, 1980); soil of sugar beet crop ( Breslin, 1979); soil under birch, pine and mixed braodleaved and coniferous stands (unpublished data); Salix ditch at seashore ( Purvis, 1982); soil under oak ( Arroyo et al., 2012, 2013a).

Counties: Carlow, Dublin, Kerry, Kildare, Wexford.

• Lysigamasus nasellus (Karg, 1968)

• Original name: Pergamasus nasellus Karg, 1968 .

• In: Deutsche Entomologische Zeitschrift, 15: 338.

Irish Records: Pasture ( Piotrowska, 2009).

Counties: Wexford.

• Lysigamasus parrunciger ( Bhattacharyya, 1963) • Original name: Pergamasus (Paragamasus) parrunciger Bhattacharyya, 1963 . • In: Bulletin of the British Museum (Natural History) Zoology, 11: 216. • Combinations/variations used in Irish literature: Pergamasus parrunciger Bhattacharyya ( Curry, 1969a, b; O’Connell, 1994), Paragamasus parrunciger Bhattacharyya (Bolger, 1980) .

Irish Records: Pasture ( Curry, 1969a, b; Bolger, 1980); fungal sporocarp ( O’Connell, 1994); soil under

Lawson cypress (unpublished data).

Counties: Kildare, Wexford.

• Lysigamasus processiferus ( Halbert, 1915) • Original name: Gamasus (Pergamasus) processiferus Halbert, 1915 • In:. Proceedings of the Royal Irish Academy, 31: 54. • Combinations/variations used in Irish literature: Gamasus (Pergamasus) processiferus Halbert ( Halbert,

1915; Hull, 1918).

Irish Records: Amongst moss ( Halbert, 1915); pasture ( Piotrowska, 2009).

Counties: Mayo, Wexford.

• Lysigamasus puerilis (Karg, 1963)

• Original name: Pergamasus puerilis Karg, 1963 .

• In: Deutsche Entomologische Zeitschrift, 10: 74.

Irish Records: Soil under conifers (unpublished data).

Counties: Dublin.

• Lysigamasus runcatellus (Berlese, 1903)

• Original name: Gamasus (Pergamasus) runcatellus Berlese, 1903

• In: Redia , 1: 237.

• Combinations/variations used in Irish literature: Gamasus (Pergamasus) runcatellus Berlese ( Halbert,

1915; Hull, 1918), Paragamasus runcatellus (Berlese) ( Purvis, 1982) , Paragamasus teutonicus Willmann

( Longworth, 1973; Curry, 1976a, 1979; Bolger, 1980; Kelly & Curry, 1985; Whelan 1986; Curry & Momen,

1988; Curry et al., 1989), Pergamasus teutonicus Willmann ( Cunningham, 1975; Ganley, 1976; O’Neill, 1981;

Purvis, 1978; Tuohy, 1979; Purvis & Curry, 1980, 1981; O’Connell, 1994), Pergamaus (Paragamasus) teutoni-

cus Willmann ( Whelan, 1976, 1978, 1985).

Irish Records: Amongst moss ( Halbert, 1915); pasture ( Cunningham, 1975; Curry, 1976a; Ganley, 1976; Purvis, 1978; Tuohy, 1979; Purvis & Curry, 1980; Bolger, 1980; O’Neill, 1981; Piotrowska, 2009); pasture on cutaway peat ( Whelan, 1976, 1978, 1985, 1986; Curry & Momen, 1988; Curry et al., 1989); manure on soil surface (Curry, 1979); Sphagnum , heather, pine shelter belt and pasture ( Longworth, 1973); dune, pasture, Iris bog, bracken bed and Salix ditch at seashore ( Purvis, 1982); winter wheat crop ( Kelly & Curry, 1985); fungal sporocarp ( O’Connell, 1994); soil under Norway spruce ( McCarthy, 2004).

Counties: Cork, Dublin, Galway, Kildare, Mayo, Meath, Offaly, Wexford.

• Lysigamasus runciger (Berlese, 1904)

• Original name: Gamasus (Pergamasus) runciger Berlese, 1904 .

• In: Redia , 1: 263.

• Combinations/variations used in Irish literature: Pergamasus runciger Berlese ( Hull, 1918; Valarasan, 1971; Cunningham, 1975; Curry, 1976a, b; Ganley, 1976; Purvis, 1978; Tuohy, 1979; Curry, 1979; Curry & O’Neill, 1979; O’Neill, 1981; Bolger & Curry, 1984; Heneghan, 1994; O’Connell, 1994; O’Connell & Bolger, 1997b), Paragamasus runciger Berlese ( Curry & O’Neill, 1979; Bolger, 1980; Purvis, 1982; Skorupski & Luxton, 1998).

Irish Records: Pasture ( Valarasan, 1971; Cunningham, 1975; Curry, 1976a, b; Ganley, 1976; Purvis, 1978; Curry & O’Neill, 1979; Tuohy, 1979; Bolger, 1980; Bolger & Curry, 1984; O’Neill, 1981; Piotrowska, 2009); manure on soil surface (Curry, 1979); Iris bog at seashore ( Purvis, 1982); fungal sporocarp ( O’Connell, 1994; O’Connell & Bolger, 1997b); soil under Norway spruce ( Heneghan, 1994); pasture and coniferous forest soils ( Arroyo et al., 2009); moss on soil, moss on tree and in soil under Sitka spruce ( Arroyo et al., 2010, 2012, 2013a); nest of Lasius flavus and pasture ( Arroyo et al., 2015).

Counties: Cavan, Cork, Dublin, Galway, Kilkenny, Kildare, Laois, Limerick, Meath, Sligo, Tipperary, Wexford, Wicklow.

• Lysigamasus schweizeri Bhattacharyya, 1963

• Original name: Pergamasus (Paragamasus) schweizeri Bhattacharyya, 1963 . • In: Bulletin of the British Museum (Natural History) Zoology, 11: 225.

• Combinations/variations used in Irish literature: Pergamasus schweizeri Bhattacharyya ( Heneghan, 1994) . Irish Records: Soil under Norway spruce ( Heneghan, 1994). Counties: Kilkenny.

• Lysigamasus truncus (Schweizer, 1961)

• Original name: Pergamasus truncus Schweizer, 1961 .

• In: Denkschriften der Schweizerischen Naturforschenden Gesellschaft, 84: 44.

• Combinations/variations used in Irish literature: Pergamasus (Paragamasus) truncus Schweizer ( Longworth, 1973; Cusack, 1974; Cusack et al., 1975; Heneghan, 1994; Skorupski & Luxton, 1998). Irish Records: Pine shelter belt on bog ( Longworth, 1973); farm grain store ( Cusack, 1974; Cusack et al., 1975); fungal fruiting body (unpublished data); soil under Norway spruce ( Heneghan, 1994; unpublished data); pasture ( Piotrowska, 2009); soil under Leyand cypress, Monterey pine, Sitka spruce and pasture (unpublished data). Counties: Kilkenny, Mayo, Wexford, Wicklow.

• Lysigamasus vagabundus (Karg, 1968)

• Original name: Pergamasus vagabundes [sic] Karg, 1968.

• In: Deutsche Entomologische Zeitschrift, 15: 342.

Irish Records: Arable soil ( Arroyo et al., 2009).

Counties: Kilkenny.

• Lysigamasus wasmanni (Oudemans, 1902) • Original name: Parasitus wasmanni Oudemans, 1902 . • In: Tijdschrift voor entomologie, 45: 39. • Combinations/variations used in Irish literature: Pergamasus wasmanni (Oudemans) ( Heneghan, 1994) .

Irish Records: Soil under Norway spruce ( Heneghan, 1994).

Counties: Kilkenny.

Neogamasus Tichomirov, 1969 • Neogamasus insignis (Holzmann, 1969) • Original name: Eugamasus insignis Holzmann, 1969 . • In: Acarologie. Schriftenreihe für Vergleichende Milbenkunde, 13: 20. • Combinations/variations used in Irish literature: Parasitus (Neogamasus) insignis (Holzman) ( Piotrowska,

2009).

Irish Records: Pasture ( Piotrowska, 2009).

Counties: Wexford.

Paragamasus Hull, 1918 • Paragamasus alpestris (Berlese, 1904) • Original name: Gamasus (Pergamasus) alpestris Berlese, 1904 . • In: Redia , 1: 236. • Combinations/variations used in Irish literature: Gamasus (Pergamasus) alpestris Berlese ( Halbert, 1915) , Pergamasus alpestris Berlese ( Hull, 1918, Valarasan, 1971, Heneghan, 1994), Pergamasus (Paragamasus) alpestris Berlese ( Whelan, 1976) .

Irish Records: Moss ( Halbert, 1915); pasture ( Valarasan, 1971); Sphagnum on bog ( Longworth, 1973);

pasture on peat ( Whelan, 1976); bracken bed at seashore ( Purvis, 1982); fungal sporocarp ( O’Connell,

1994); soil under Norway spruce ( Heneghan, 1994); moss on Sitka spruce ( Arroyo et al., 2010).

Counties: Dublin, Kilkenny, Laois, Mayo, Wexford.

• Paragamasus diversus ( Halbert, 1915)

• Original name: Gamasus (Pergamasus) diversus Halbert, 1915 .

• In: Proceedings of the Royal Irish Academy, 31: 53.

• Combinations/variations used in Irish literature: Gamasus diversus Halbert (Halbert, 1915) , Pergamasus diversus Halbert ( Hull, 1918, Purvis, 1978, O’Connell, 1994).

Irish Records: Under stone at lakeside ( Halbert, 1915); under pines and rhododendrons ( Bhattacharyya, 1963); heather on bog ( Longworth, 1973); pasture ( Purvis, 1978; unpublished data); winter wheat crop ( Kelly & Curry, 1985); nest of Lasius flavus and pasture ( Arroyo et al., 2015); mixed broadleaf and ash woodlands (unpublished data); fungal sporocarp ( O’Connell, 1994); soil in winter wheat field (unpublished data); moss on soil under oak and Sitka spruce; soil under ash; moss on oak and Sitka spruce ( Arroyo & Bolger, 2007; Arroyo et al., 2010, 2012, 2013a).

Counties: Carlow, Cork, Dublin, Galway, Kerry, Kildare, Laois, Limerick, Mayo, Sligo.

• Paragamasus integer ( Bhattacharyya, 1963) • Original name: Pergamasus (Paragamasus) integer Bhattacharyya, 1963 . • In: Bulletin of the British Museum (Natural History) Zoology, 11: 197. • Combinations/variations used in Irish literature: Pergamasus integer Bhattacharyya ( O’Connell, 1994) .

Irish Records: Fungal sporocarp ( O’Connell, 1994; unpublished data).

Counties: Wicklow.

• Paragamasus robustus (Oudemans, 1902)

• Original name: Parasitus longulus var. robustus Oudemans, 1902 .

• In: Tijdschrift voor entomologie, 45: 38.

• Combinations/variations used in Irish literature: Gamasus (Pergamasus) robustus Oudemans ( Halbert,

1915), Pergamasus robustus (Oudemans) ( Hull, 1918; Valarasan, 1971; Heneghan, 1994; O’Connell, 1994;

O’Connell & Bolger, 1997b).

Irish Records: In moss and under pine needles ( Halbert, 1915); The Bills Rock ( Halbert, 1915); heather, pine shelter belt and pasture on bog ( Longworth, 1973); soil under oak, pine, Sitka spruce and mixed stands of broadleaves and conifers (unpublished data); pasture ( Valarasan, 1971; unpublished data); dune at seashore ( Purvis, 1982); fungal sporocarp ( O’Connell, 1994; O’Connell & Bolger, 1997b; unpublished data); soil under Norway spruce ( Heneghan, 1994); ash woodland (unpublished data); pasture, rough grazing, coniferous forest and arable soils ( Arroyo et al., 2009); small mammal nests (unpublished data); moss on soil under oak and Scots pine; soil under oak, Scots pine and Sitka spruce; moss on Scots pine ( Arroyo & Bolger, 2007; Arroyo et al., 2010, 2012, 2013a); woodland, cliff and woodland (Bolger, in press); fen, Atlantic blanket bog, raised bog ( Wisdom, 2010; Wisdom et al., 2011).

Counties: Cork, Dublin, Galway, Kerry, Kildare, Kilkenny, Laois, Mayo, Offaly, Roscommon, Waterford, Wexford, Wicklow.

• Paragamasus rothamstedensis ( Bhattacharyya, 1963)

• Original name: Pergamasus (Paragamasus) rothamstedensis Bhattacharyya, 1963 .

• In: Bulletin of the British Museum (Natural History) Zoology, 11: 219.

• Combinations/variations used in Irish literature: Paragamasus rothamstedensis Bhattacharyya ( Piotrowska, 2009) . Irish Records: Pasture ( Piotrowska, 2009). Counties: Wexford.

• Paragamasus similis (Willmann, 1953)

• Original name: Pergamasus similis Willmann, 1953 .

• In: Denkschriften der Kaiserlichen Akademie der Wissenschaften / Mathematisch -Naturwissenschaftliche Classe - I, 162: 449. Irish Records: Pasture, rough grazing, broadleaved and coniferous forest ( Arroyo et al., 2009); moss on soil under oak, and Sitka spruce; soil under oak; moss on oak; canopy of oak ( Arroyo et al., 2012, 2013a, b); pasture ( Piotrowska, 2009); raised bog ( Wisdom, 2010; Wisdom et al., 2011). Counties: Cork, Galway, Kerry, Laois, Meath, Roscommon, Sligo, Tipperary, Waterford, Wicklow.

Parasitellus Willmann, 1939

• Parasitellus fucorum (De Geer, 1778)

• Original name: Acarus fucorum De Geer, 1778 .

• In: Memoires pour servir a I’histoire des insectes. Stockholm, 7: 112.

Irish Records: On Psithyrus rupestris (Halbert unpublished notes); on Ottiorynchus sulcatus from a carpet ( Hyatt, 1990b).

Counties: Galway, Wicklow.

• Parasitellus ignotus Vitzhum, 1930

• Original name: Parasitus ignotus Vitzthum, 1930 .

• In: Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere, 60: 40.

Irish Records: In the burrow of a sand martin ( Riparia riparia ) ( Hyatt, 1980).

Counties: Cork.

• Parasitellus talparum (Oudemans, 1913) • Original name: Parasitus talparum Oudemans, 1913 . • In: Entomologische Berichten, 3: 333. • Combinations/variations used in Irish literature: Parasitus talparum Oudemans ( Turk & Turk, 1952).

Irish Records: On Apodemus sylvaticus ( Turk & Turk, 1952).

Counties: Dublin.

Parasitus Latreille, 1795

• Parasitus americanus (Berlese, 1905)

• Original name: Gamasus americanus Berlese, 1905 .

• In: Redia , 3: 138.

Irish Records: Mushroom compost (Hyatt, 1988).

Counties: Kildare.

• Parasitus beta Oudemans & Voigts, 1904 • Original name: Parasitus beta Oudemans & Voigts, 1904 . • In: Zoologischer Anzeiger, 27: 652. • Combinations/variations used in Irish literature: Parasitus eta Oudemans & Voigts ( Ganley, 1976; Whelan, 1976; Purvis, 1978; Curry & O’Neill, 1979; Bolger, 1980; Purvis & Curry, 1980).

Irish Records: Pasture ( Ganley, 1976; Purvis, 1978; Curry & O’Neill, 1979; Purvis & Curry, 1980; Bolger,

1980; Piotrowska, 2009); pasture on peat ( Whelan, 1976).

Counties: Dublin, Galway, Kildare, Meath, Westmeath, Wexford.

• Parasitus coleoptratorum (Linnaeus, 1758)

• Original name: Acarus coleoptratorum Linnaeus, 1758 .

• In: Systema Naturae 10th edn. Stockholm (Salvii). 1, 618.

• Combinations/variations used in Irish literature: Gamasus (Gamasus) coleoptratorum (Linnaeus) ( Halbert,

1915).

Irish Records: Under bark of decaying trees ( Halbert, 1915); under stones resting on sand, gravel, and shells on upeer seashore ( Halbert, 1915, 1920); manure on soil surface (Curry, 1979); pasture ( Ganley, 1976; Purvis, 1978; Bolger, 1980; O’Neill, 1981); in compost, on Geotrupes stercorarius , in nest of Starling ( Sturnus vulgaris ) and burrow of sand martin ( Riparia riparia ) ( Hyatt, 1980; unpublished data).

Counties: Clare, Cork, Dublin, Kildare, Mayo, Meath, Sligo, Westmeath, Wexford.

• Parasitus consanguineus Oudemans & Voigts, 1904

• Original name: Parasitus consanguineus Oudemans & Voigts, 1904 .

• In: Zoologischer Anzeiger, 27: 651.

Irish Records: Starling’s nest ( Sturnus vulgaris ) Hyatt, 1980); pasture (Bolger, 1980; O’Neill, 1981); rotting vegetable matter ( Hyatt, 1990a).

Counties: Clare, Cork, Meath, Wexford.

• Parasitus evertsi Oudemans, 1902

• Original name: Parasitus evertsi Oudemans, 1902 .

• In: Nederlandsch Tijdschrift voor de Dierkunde, 1: 283.

Irish Records: Flood debris ( Hyatt, 1980; unpublished data); fungal sporocarp ( O’Connell, 1994).

Counties: Leitrim.

• Parasitus fimetorum (Berlese, 1904)

• Original name: Gamasus fimetorum Berlese, 1904 .

• In: Redia , 1: 238.

• Combinations/variations used in Irish literature: Gamasus (Gamasus) fimetorum Berlese ( Halbert, 1915;

Hull, 1918), Parasitus hibernicus Turk & Turk ( Turk & Turk, 1952).

Irish Records: Under bark ( Halbert, 1915); nest of kittiwake ( Rissa tridactyla ) ( Turk & Turk, 1952); pasture ( Curry, 1969a; Purvis, 1978; Bolger, 1980); manure on soil surface (Curry, 1979); in nest of Starling ( Sturnus vulgaris ) and burrow of Sand Martin ( Riparia riparia ) and moss on trees ( Hyatt, 1980 and unpublished data); dune at seashore ( Purvis, 1982).

Counties: Clare, Cork, Kildare, Mayo, Meath, Westmeath. Wexford.

• Parasitus hyalinus (Willmann, 1949)

• Original name: Eugamasus hyalinus Willmann, 1949 .

• In: Veroeffentlichungen aus dem Museum für Naturkunde Bremen, Al: 110.

Irish Records: Pasture ( Hyatt, 1980; Bolger, 1980; Bolger & Curry; 1984) pasture on cutaway peat ( Curry & Momen, 1988).

Counties: Offaly, Wexford.

• Parasitus kempersi Oudemans, 1902

• Original name: Gamasus kempersi Oudemans, 1902 .

• In: Tijdschrift voor entomologie, 45: 36.

• Combinations/variations used in Irish literature: Gamasus (Gamasus) Kempersi Oudemans ( Halbert, 1915,

1920; Hull, 1918).

Irish Records: Under stones and seaweed and amongst moist, shelly sand and gravel on upper levels of seashore ( Halbert, 1915, 1920); rotting seaweed and linestone pavement ( Hyatt, 1980; unpublished data); dry tidal debris and Armeria tussocks (Healy & McGrath, 1988).

Counties: Clare, Cork, Dublin, Mayo, Wexford.

• Parasitus loricatus (Wankel, 1861)

• Original name: Gamasus loricatus Wankel, 1861 .

• In: Sonderabdruck der Sitzungsberichte der kais. Akademie der Wissenschaften in Wien, 43: 261.

• Combinations/variations used in Irish literature: Eugamasus loricatus Wankel ( O’Farrell & Butler, 1948) .

Irish Records: Stored foodstuffs ( O’Farrell & Butler, 1948); small mammal nests (unpublished data); in nest of Starling ( Sturnus vulgaris ) and burrow of Sand Martin ( Riparia riparia ) and tree litter around roadway (Hyatt; unpublished data).

Counties: Antrim, Cork, Derry, Westmeath.

• Parasitus lunulatus (J. Müller, 1859)

• Original name: Porrhostaspis lunulata Müller, 1859 .

• In: Lotos 9: 29.

• Combinations/variations used in Irish literature: Gamasus (Eugamasus) cornutus G. & R. Canestrini ( Halbert, 1915; Hull, 1918), Eugamasus cornutus (Canestrini) ( Curry, 1969a; Valarasan, 1971; Longworth, 1973), Eugamasus lunulatus (Müller) ( Whelan, 1976) . Irish Records: Under bark ( Halbert, 1915); on seashore (Halbert unpublished notes); pasture ( Curry, 1969a; Valarasan, 1971); pasture on peat ( Whelan, 1976); soil in winter wheat field (unpublished data); pine shelter belt and pasture on bog ( Longworth, 1973); leaf litter ( Hyatt, 1990a); fungal sporocarp ( O’Connell, 1994); soil under horse chestnut, oak and mixed broadleaves (unpublished data). Counties: Carlow, Clare, Dublin, Kerry, Kildare, Kilkenny, Mayo, Sligo.

• Parasitus nolli (Karg, 1965)

• Original name: Eugamasus nolli Karg, 1965 .

• In: Mitteilungen aus dem Zoologischen Museum in Berlin, 41: 318.

Irish Records: Pasture ( Ganley, 1976).

Counties: Dublin.

Pergamasus Berlese, 1904 • Pergamasus alpinus (Berlese, 1903) • Original name: Gamasus (Pergamasus) theseus var. alpinus Berlese, 1903 . • In: Redia , 1: 236. • Combinations/variations used in Irish literature: Pergamasus alpestris Berlese var. alpinus Berlese ( Hull,

1918).

Irish Records: In moss ( Hull, 1918).

Counties: Not specified.

• Pergamasus crassipes (Linnaeus, 1758) Berlese, 1906

• Original name: Acarus crassipes Linnaeus, 1758 .

• In: Redia , 3: 229 (Male) Redia 3: 232 (Female).

Irish Records: Common in moss and under bark ( Halbert, 1915); pasture ( Curry, 1976a; Valarasan, 1971; Ganley, 1976; Piotrowska, 2009); pine shelter bed on bog ( Longworth, 1973); pasture on peat ( Whelan, 1976); soil in winter wheta field (unpublished data); soil under mixed broadleaved and conferous stands (unpublished data); pasture, broadleaved forest and arable soil ( Arroyo et al., 2009); nest of Lasius flavus and pasture ( Arroyo et al., 2015); rough grassland, heather, cliff, moss (Bolger, in press); under Western hemlock and Norway spruce (unpublished data); fungal sporocarp ( O’Connell, 1994; O’Connell & Bolger, 1997b); soil under Norway spruce ( Heneghan, 1994; unpublished data); moss under oak, ash, Scots pine and Sitka spruce; soil under oak, Scots pine and Sitka spruce; moss on Scots pine, yew and Sitka spruce; canopy of ash, oak and Sitka spruce ( Arroyo & Bolger, 2007; Arroyo et al., 2010, 2012, 2013a, b); fixed dunes of Machair system ( Arroyo & Bolger, 2011); riverbank (unpublished data); raised bog, Atlantic blanket bog and fen ( Wisdom, 2010; Wisdom et al., 2011).

Counties: Clare, Cork, Donegal, Dublin, Galway, Kerry, Kildare, Kilkenny, Laois, Limerick, Louth, Mayo, Offaly, Roscommon, Sligo, Tipperary, Waterford, Wexford, Wicklow.

• Pergamasus hortensis Bhattacharyya, 1963

• Original name: Pergamasus hortensis Bhattacharyya, 1963 .

• In: Bulletin of the British Museum (Natural History) Zoology, 11: 173.

Irish Records: Pasture ( Curry, 1969a).

Counties: Kildare.

• Pergamasus longicornis ( Berlese, 1906)

• Original name: Gamasus (Pergamasus) crassipes var. l ongicornis Berlese, 1906.

• In: Redia , 3: 232.

• Combinations/variations used in Irish literature: Gamasus crassipes var longicornis Berlese ( Halbert,

1920).

Irish Records: Under stones and decaying seaweeds ( Halbert, 1920); moss on rock on lakeshore ( Bhattacharyya, 1963); pasture ( Curry, 1969a, 1973, 1976a; Valarasan, 1971; Cunningham, 1975; Ganley, 1976; Purvis, 1978; Curry & O’Neill, 1979; Tuohy, 1979; Purvis & Curry, 1980, 1981; Bolger, 1980; O’Neill, 1981; Piotrowska, 2009); manure on soil surface (Curry, 1979); soil of sugar beet crop ( Breslin, 1979); pine shelter bed on bog ( Longworth, 1973); soil in saltmarsh ( Healy, 1975; Speight & Healy, 1977); soil in barley and winter wheat fields ( Emmanuel, 1977) unpublished data); dune and Iris bog at seashore ( Purvis, 1982); winter wheat crop ( Kelly & Curry, 1985); soil under Norway spruce ( Heneghan, 1994; McCarthy, 2004); small mammal nests (unpublished data); arable soil and cliffs (Bolger, in press); fungal sporocarp ( O’Connell, 1994; O’Connell & Bolger, 1997b, unpublished data); soil under pasture and Sitka spruce (unpublished data).

Counties: Carlow, Clare, Dublin, Galway, Kildare, Kilkenny, Mayo, Meath, Wexford, Wicklow.

• Pergamasus norvegicus (Berlese, 1905) • Original name: Gamasus septentrionalis var. norvegicus Berlese, 1905 . • In: Redia , 3, 190. • Combinations/variations used in Irish literature: Gamasus (Amblygamasus) septentrionalis var. norvegicus Berlese ( Halbert, 1915) .

Irish Records: In moss ( Halbert, 1915); unattributed habitat Kilruddery (Halbert unpublished notes)

Counties: Mayo, Wicklow.

• Pergamasus noster (Berlese, 1903)

• Original name: Gamasus (Pergamasus) noster Berlese, 1903 .

• In: Redia , 1: 236.

Irish Records: Fungal sporocarp ( O’Connell, 1994).

Counties: Kilkenny.

• Pergamasus quisquiliarum (G. & R. Canestrini, 1882)

• Original name: Gamasus quisquiliarum Canestrini, G. & R., 1882.

• In: Atti del Reale Istituto veneto di scienze, lettere ed arti, Series 5, 8: 920.

Irish Records: Pasture ( Valarasan, 1971; Ganley, 1976; Bolger, 1980; Piotrowska, 2009); fungal sporocarp ( O’Connell, 1994); soil in winter wheta field (unpublished data).

Counties: Carlow, Cork, Dublin, Galway, Meath, Wexford.

• Pergamasus septentrionalis (Oudemans, 1902)

• Original name: Parasitus septentrionalis Oudemans, 1902 .

• In: Tijdschrift voor entomologie, 45: 39.

Irish Records: Moss on rock ( Bhattacharyya, 1963); pasture ( Curry, 1969a; Valarasan, 1971; Bolger, 1980; Piotrowska, 2009); manure on soil surface (Curry, 1979); pine shelter belt and pasture on bog ( Longworth, 1973); soil in barley field ( Emmanuel, 1977); soil under oak (unpublished data); cliffs, ant nest, rough grassland (Bolger, in press); fungal sporocarp ( O’Connell, 1994; unpublished data); soil under Leyland cypress, Sitka spruce and pasture (unpublished data); moss on oak and Sitka spruce; moss on soil under oak, ash and Sitka spruce; soil under ash and Sitka spruce; canopy of Sitka spruce ( Arroyo & Bolger, 2007; Arroyo et al., 2010, 2012, 2013a, b); fore dunes of Machair system ( Arroyo & Bolger, 2011); mountain blanket bog ( Wisdom, 2010, Wisdom et al., 2011).

Counties: Clare, Donegal, Dublin, Galway, Kerry, Kildare, Kilkenny, Laois, Mayo, Meath, Offaly, Roscommon, Wexford, Wicklow.

Poecilochirus G. & R Canestrini, 1882

• Poecilochirus carabi G.and R. Canestrini, 1882

• Original name: Poecilochirus carabi G.and R. Canestrini, 1882.

• In: Atti del Reale Istituto veneto di scienze, lettere ed arti, Series 5, 8: 56.

• Combinations/variations used in Irish literature: Gamasoides carabi G.and R. Canestrini ( Halbert, 1915); Poecilochirus necrophori Vitzthum ( Turk & Turk, 1952; Douthwaite, 1966). Irish Records: On the carabid Pterostichus niger ( Halbert, 1915) ; on Apodemus sylvaticus ( Douthwaite, 1966) ; on the burying beetles, Nicrophorus humator and N. investigator , in rat carcasses ( Sleeman, 1983); from Nicrophagus investigator ( Turk & Turk, 1952; Hyatt, 1990a, b). Counties: Cork, Kerry, Wexford, Wicklow.

• Poecilochirus davydovae Hyatt, 1980

• Original name: Poecilochirus davydovae Hyatt, 1980 .

• In: Bulletin of the British Museum (Natural History) Zoology, 38: 358.

Irish Records: On Nicrophagus investigator ( Hyatt, 1990a) .

Counties: Wexford.

• Poecilochirus subterraneus (Müller, 1859) • Original name: Porrhostaspis subterranea Müller, 1859 . • In: Jahresheft der Naturwissenschaftlieben Section der k. k. magr. schles. Gesellschaft fur Ackerbau, Nature -

und Landeskunde. 1859: 176.

Irish Records: On Nicrophagus investigator ( Turk & Turk, 1952; Hyatt, 1990a, b).

Counties: Wexford.

Trachygamasus Berlese, 1906

• Trachygamasus ambulacralis (Willmann, 1949)

• Original name: Saprogamasus ambulacralis Willmann, 1949 .

• In: Veröffentlichungen aus dem Museum für Natur -, Völker - und Handelskunde Bremen, Reihe A, Naturwissenschaften, 1: 137. Irish Records: Seaweed ( Hyatt, 1980). Counties: Clare.

• Trachygamasus gracilis Karg, 1965

• Original name: Saprogamasus gracilis Karg, 1965 .

• In: Mitteilungen aus dem Zoologischen Museum in Berlin, 41: 300.

Irish Records: Pasture ( Purvis, 1978; Bolger, 1980; Bolger & Curry, 1980; Hyatt, 1980); manure on soil surface (Curry, 1979).

Counties: Kildare, Meath, Wexford.

Vulgarogamasus Tichomirov, 1969 • Vulgarogamasus burchanensis (Oudemans, 1903) • Original name: Parasitus burchanensis Oudemans, 1903 . • In: Entomologische Berichten, 1: 86. • Combinations/variations used in Irish literature: Eugamasus butleri ( O’Farrell & Butler, 1948; Cusack, 1974; Cusack et al., 1975; Hughes. 1976).

Irish Records: Farm grain stores, flour mills and railway wagons which contained grain ( Hughes, 1948;

O’Farrell & Butler, 1948; Evans, 1958; Cusack, 1974; Cusack et al., 1975; Hyatt, 1980).

Counties: Antrim, Derry.

• Vulgarogamasus immanis (Berlese, 1904)

• Original name: Gamasus (Eugamasus) immanis Berlese, 1904 .

• In: Redia , 1: 262.

• Combinations/variations used in Irish literature: Gamasus (Eugamasus) immanis Berlese ( Halbert, 1915; Hull, 1918).

Irish Records: Under stones, amongst shingle, and in decaying seaweed at high-water mark on seashore ( Berlese 1906; Halbert, 1915, 1920).

Counties: Cork, Dublin, Galway, Mayo.

• Vulgarogamasus kraepelini (Berlese, 1905)

• Original name: Gamasus (Eugamasus) kraepelini Berlese, 1905 .

• In: Redia , 2: 232.

• Combinations/variations used in Irish literature: Gamasus (Eugamasus) magnus Berlese ( Halbert, 1915; Hull, 1918), Eugamasus kraepelini Berlese ( Curry, 1969a, b; Valarasan, 1971; Purvis, 1978), Parasitus kraepelini (Berlese) ( Ganley, 1976) . Irish Records: Moss in woodland ( Halbert, 1915); on decaying fungi in woodland ( Halbert, 1915); pasture ( Curry, 1969a, b; Valarasan, 1971; Ganley, 1976; Purvis, 1978); fungal sporocarp ( O’Connell, 1994; unpublished data). Counties: Dublin, Kildare, Mayo, Westmeath, Wicklow.

• Vulgarogamasus oudemansi (Berlese, 1904)

• Original name: Gamasus (Eugamasus) oudemansi Berlese, 1904 .

• In: Redia , 1: 280.

Irish Records: Hyatt, 1980.

Counties: Cork.

• Vulgarogamasus trouessarti (Berlese, 1892) • Original name: Gamasus trouessarti Berlese, 1892 . • In: Acari ., Myriapoda and Scorpiones hucusque in Italia reperta, 3: 67. • Combinations/variations used in Irish literature: Gamasus trouessarti (Berlese) ( Halbert, 1915, 1920).

Irish Records: Variety of habitats on seashore ( Halbert, 1915, 1920).

Counties: Dublin, Galway, Mayo.