Lindsaea fuscopetiolata A.Rojas & Tejero

MONRO, A. K., SANTAMARÍA-AGUILAR, D., GONZÁLEZ, F., CHACÓN, O., SOLANO, D., RODRÍGUEZ, A., ZAMORA, N., FEDELE, E. & CORREA, M., 2017, A first checklist to the vascular plants of La Amistad International Park (PILA), Costa Rica-Panama, Phytotaxa 322 (1), pp. 448-450 : 448-450

Lindsaea fuscopetiolata A.Rojas & Tejero Danaea moritziana C.Presl

Terrestrial herbs. Distribution in PILA 1600–1700 Terrestrial herbs. Distribution in PILA 200–1700 m, low m, mixed forest in transition to cloud forest. Global elevation mixed forest, mixed forest in transition to low distribution: Mexico, Honduras, Costa Rica, Panama. elevation forest, mixed forest in transition to cloud forest. Records: D. Santamaría 4411 (INB). Uses: medicinal. Global distribution: Costa Rica to Venezuela, French Guyana, Bolivia. Records: A. Rodríguez 11508 (BM, Lindsaea imrayana (Hook.) Pérez Arbel. INB, MO, PMA ).

Facultatively epiphytic herbs. Distribution in PILA

1100–1800 m, mixed forest in transition to low elevation Eupodium pittieri (Maxon) Christenh.

forest, mixed forest in transition to cloud forest. Global Terrestrial herbs. Distribution in PILA 1500–1700 distribution: Costa Rica to Venezuela, the Guyanas, m, mixed forest in transition to cloud forest. Global Greater Antilles, Lesser Antilles. Records: A. Rodríguez distribution: Costa Rica to Bolivia and Venezuela, 10648 (BM, INB, MO, PMA), D. Santamaría 5758 (BM, Trinidad (see Christenhusz 2010). Records: A. Fernández INB, MO, PMA), D. Solano 3924 (BM, INB, MO, PMA), 690 (INB), M. Grayum 10310 (INB), M. Grayum 10379 G. Herrera 2667 (INB), G. Herrera 2667 (MO), G. (INB).

Herrera 5388 (INB), G. Herrera 6190 (CR, INB, MO), J.

Bittner 1480 (MO), M. Grayum 10368 (CR, INB, MO). Marattia excavata Underw.

Terrestrial herbs. Distribution in PILA 1000–2200 m, mixed forest in transition to low elevation forest, mixed Alansmia senilis (Fée) Moguel & M.Kessler forest in transition to cloud forest, cloud forest, low Epiphytic herbs. Distribution in PILA 1900–3300 m, elevation oak forest. Global distribution: Mexico to cloud forest, low elevation oak forest, páramo. Global Panama. Records: G. Herrera 5271 (MO). distribution: Mexico to Bolivia. Records: A. Chacón 544 (INB), A. Rojas 3196 (CR, INB), G. Davidse 25469

(MO), G. Davidse 25789 (MO), G. Davidse 28665 (MO),

OPHIOGLOSSACEAE G. Davidse 28872 (MO), G. Davidse 28913 (MO), G. Davidse 29226 (MO), G. Davidse 29242A (MO), G.

Botrychium virginianum (L.) Sw. Herrera 3679 (INB), G. Herrera 3716 (INB, MO), G. Terrestrial herbs. Distribution in PILA 1500–3000 m, Herrera 6160 (INB), F. Quesada 1571 (INB), M. Moraga mixed forest in transition to cloud forest, cloud forest, 317 (CR, INB). low elevation oak forest, high elevation oak forest. Global distribution: Canada to Mexico, Guatemala, Eurasia. Alansmia turrialbae (Christ) Moguel & M.Kessler Records : F. Quesada 1600 (INB). Epiphytic herbs. Distribution in PILA 1000–3100 m, mixed forest in transition to low elevation forest, mixed

Ophioglossum palmatum L. forest in transition to cloud forest, cloud forest, low Epiphytic herbs. Distribution in PILA 1400–2600 m, elevation oak forest, high elevation oak forest, páramo. mixed forest in transition to cloud forest, cloud forest, low Global distribution: Costa Rica, Panama. Records: A. elevation oak forest, high elevation oak forest, pasture. Chacón 502 (MO), G. Herrera 3642 (INB), G. Davidse Global distribution: USA to Mexico to Suriname, Greater 28892 (CR, MO), G. Davidse 29145 (CR, MO). Antilles, Lesser Antilles. Records: A. Fernández 740 (INB, MO), A. Fernández 828 (INB, MO), A. Rodríguez Arachniodes denticulata (Sw.) Ching 10649 (BM, INB, MO, PMA), A. Soto 303 (INB), D. Terrestrial herbs. Distribution in PILA 0–3300 m, low Solano 3829 (BM, INB, MO, PMA), D. Solano 4192 elevation mixed forest, mixed forest in transition to low (BM, INB, MO, PMA), E. Alfaro 3674 (INB), G. Davidse elevation forest, mixed forest in transition to cloud forest, 28822 (MO), G. Herrera 5905 (INB, MO), G. Herrera cloud forest, low elevation oak forest, páramo. Global 5989 (CR, INB, MO), G. Herrera 6185 (CR, INB, MO), distribution: Mexico to Bolivia, Greater Antilles. Records: F. Quesada 1427 (INB), F. Quesada 1570 (INB). A. Chacón 538 (INB, MO), A. Fernández 957 (CR, INB), A.K. Monro 4124 (BM, INB, MO), A.K. Monro 4159

Ophioglossum reticulatum L. (BM), A. Rodríguez 10895 (BM, INB, MO, PMA), A. Terrestrial herbs. Distribution in PILA 0–1800 m, low Rodríguez 11834 (BM, INB, MO, PMA), A. Rojas 3251 elevation mixed forest, mixed forest in transition to low (CR, INB), D. Santamaría 5760 (BM, INB, MO, PMA), elevation forest, mixed forest in transition to cloud forest, E. Alfaro 3336 (INB), E. Alfaro 5606 (BM, INB), F. cloud forest. Global distribution: Mexico to Argentina, Quesada 1456 (CR, INB, MO), G. Davidse 25460 (CR, Greater Antilles, Lesser Antilles, Africa, Asia. Records: F. MO), G. Davidse 28629 (CR, MO), G. Davidse 28860 Quesada 1555 (INB). (CR, MO), G. Davidse 29393 (MO), G. Davidse 23881 (MO), G. Herrera 5281 (CR, MO), G. Herrera 6134 (CR,

POLYPODIACEAE INB ), J. Bittner 1737 (INB, MO), M.A. Thomas 1579 (MO), M.A. Thomas 1619 (MO), M.A. Thomas 1629

Alansmia cultrata (Bory ex Willd.) Moguel & M.Kessler (MO) , M. Moraga 460 (CR, INB, MO), M. Valerio 165 Epiphytic herbs. Distribution in PILA 1100–3100 m, (CR, MO). mixed forest in transition to low elevation forest, mixed forest in transition to cloud forest, cloud forest, low Arachniodes ochropteroides (Baker) Lellinger elevation oak forest, high elevation oak forest, páramo. Facultatively epiphytic herbs. Distribution in PILA 700– Global distribution: Mexico to Brazil, Greater Antilles, 1600 m, low elevation mixed forest, mixed forest in Lesser Antilles. Records: A. Fernández 661 (CR, INB), transition to low elevation forest. Global distribution: A. Fernández 939 (INB), A. Fernández 966 (CR, INB), Costa Rica to Ecuador, Venezuela, Greater Antilles. D. Santamaría 5686 (BM, INB, MO, PMA), G. Davidse Records: G. Herrera 5291 (CR, INB, MO), G. Herrera 28893 (MO), G. Davidse 29013 (MO), G. Davidse 29082 5378 (CR, INB). (CR, MO), G. Davidse 29229 (MO), G. Davidse 29233A (MO), G. Herrera 5302 (CR, INB, MO), G. Herrera 6125 Campyloneurum amphostenon (Kunze ex Klotzsch) Fée (CR, INB) , J. Bittner 1791 (INB, MO), J. Bittner 1856 Facultatively epiphytic herbs. Distribution in PILA 1100– (INB, MO), F. Quesada 1437 (CR, INB, MO), F. Quesada 3500 m, mixed forest in transition to low elevation forest, 1478 (CR, INB). mixed forest in transition to cloud forest, cloud forest, low elevation oak forest, páramo. Global distribution:

Mexico to Bolivia, Greater Antilles. Records: A. Chacón Epiphytic herbs. Distribution in PILA 0–1500 m, low 550 (INB), A. Rojas 3224 (INB, MO), B.B. Klitgaard 806 elevation mixed forest, mixed forest in transition to low (BM, INB, MO), B. Gamboa 595,2 (INB), B. Gamboa 655 elevation forest, mixed forest in transition to cloud forest. (INB, MO), E. Alfaro 3332 (INB), G. Davidse 23965 (CR, Global distribution: Mexico to Bolivia, Greater Antilles. MO), G. Davidse 25333 (CR, MO), G. Davidse 25837 Records: E. Mora 293 (CR, INB).

(CR, MO), G. Davidse 25897 (MO), G. Davidse 26019

(MO), G. Davidse 28649 (MO), G. Davidse 28746 (MO), Campyloneurum sphenodes (Kunze & Klotzsch) Fée. G. Davidse 29070 (MO), G. Davidse 29328 (CR, MO), G. Epiphytic herbs. Distribution in PILA 100–2200 m, low Herrera 3673 (INB, MO), G. Herrera 3719 (INB, MO), elevation mixed forest, mixed forest in transition to low G. Herrera 5901 (CR, INB, MO), G. Herrera 6147 (CR, elevation forest, mixed forest in transition to cloud forest, INB), G. McPherson 15886 (MO), F. Quesada 1468 (CR, cloud forest, low elevation oak forest. Global distribution: INB), F. Quesada 1409 (CR, INB), M. Moraga 310 (CR, Costa Rica to Bolivia. Records: A. Fernández 654 INB), S. Knapp 9909 (BM, INB, MO). (CR, INB, MO), A. Fernández 725 (CR, INB, MO), A. Fernández 931 (CR, INB, MO), A. Fernández 999 (CR,

Campyloneurum angustifolium (Sw.) Fée INB, MO ), A.K. Monro 5626 (BM, INB, MO, PMA), A. Epiphytic herbs. Distribution in PILA 100–1500 m, low Rodríguez 10630 (BM, INB, MO, PMA), A. Rodríguez elevation mixed forest, mixed forest in transition to low 11228 (BM, INB, MO, PMA), B. Gamboa 654 (INB), D. elevation forest, mixed forest in transition to cloud forest. Santamaría 3817 (INB), D. Santamaría 5750 (BM, INB, Global distribution: USA to Argentina, Greater Antilles. MO, PMA), D. Santamaría 6158 (BM, INB, MO, PMA), Records: A. Rodríguez 11381 (BM, INB, MO, PMA), D. Solano 3951 (BM, INB, MO, PMA), D. Solano 4183 C. Hamilton 3875 (MO), G. Davidse 25464 (MO), G. (BM, INB, MO, PMA), E. Alfaro 775 (INB), G. Herrera Davidse 25728 (MO), G. Herrera 5174 (CR, INB, MO), 5997 (CR, INB), J. Bittner 1500 (MO), J. Bittner 1739 F. Quesada 1461 (CR, INB), M. Moraga 434 (CR, INB, (INB), J. Bittner 1838 (INB), J. Bittner 1920 (INB), F. MO). Quesada 1681 (INB), M. Moraga 498 (CR, INB), S. Bridgewater 4136 (BM, INB, MO, PMA).

Campyloneurum aphanophlebium (Kunze) T.Moore

Facultatively epiphytic herbs. Distribution in PILA 100– □ ε-PILA Ceradenia aulaeifolia L.E.Bishop. ex A.R.Sm. 600 m, low elevation mixed forest. Global distribution: Epiphytic herbs. Distribution in PILA 3000–3500 m, high Mexico to Brazil. Records: A. Rodríguez 11265 (BM, elevation oak forest, páramo. Global distribution: Costa INB, MO, PMA). Rica, Panama. Records: S. Knapp & A.K. Monro 9867 (BM, MO, PMA), S. Knapp & A.K. Monro 9956 (BM,

Campyloneurum densifolium (Hieron.) Lellinger MO, PMA ), A. Rojas 3232 (INB, MO), G. Davidse 25414 Terrestrial herbs. Distribution in PILA 2400–3300 m, low (MO), G. Davidse 25939 (CR, MO), G. Davidse 25966 elevation oak forest, páramo. Global distribution: Mexico, (MO), G. Davidse 26048 (CR, MO).

Guatemala, Costa Rica to Brazil, Greater Antilles, Lesser

Antilles. Records: G. Davidse 29003 (MO). Ceradenia fucoides (Christ) L.E.Bishop

Epiphytic herbs. Distribution in PILA 1900–2500 m,

Campyloneurum falcoideum (Kuhn ex Hieron.) M.Mey. cloud forest, low elevation oak forest. Global distribution: ex Lellinger Costa Rica, Panama. Records: A. Fernández 832 (INB), Epiphytic herbs. Distribution in PILA 1300–2700 m, A. Fernández 977 (CR, INB), A. Rodríguez 10702 (BM, mixed forest in transition to cloud forest, cloud forest, INB, MO, PMA), G. Davidse 28719 (CR, MO), G. low elevation oak forest, high elevation oak forest. Davidse 28747 (CR, MO), G. Herrera 3410 (MO), G. Global distribution: Costa Rica to Venezuela. Records: Herrera 5446 (INB), G. Herrera 5502 (INB), G. Herrera B. Gamboa 651 (INB), G. Davidse 29159 (CR, MO), M. 5884 (CR, INB, MO), G. Herrera 5977 (INB), G. Herrera Moraga 471 (CR, INB). 6127 (CR, INB, MO), J. Bittner 1802 (INB, MO), F. Quesada 1410 (CR, INB, MO), F. Quesada 1956 (INB).

Campyloneurum fasciale (Humb. & Bonpl. ex Willd.)

C.Presl Ceradenia jungermannioides (Klotzsch) L.E.Bishop Epiphytic herbs. Distribution in PILA 200–1600 m, low Epiphytic herbs. Distribution in PILA 600–3100 m, low elevation mixed forest, mixed forest in transition to low elevation mixed forest, mixed forest in transition to low elevation forest. Global distribution: Mexico to Bolivia. elevation forest, mixed forest in transition to cloud forest, Records: A. Rodríguez 11735 (BM, INB, MO, PMA), L. cloud forest, low elevation oak forest, high elevation oak Angulo 149 (CR, INB), L. Angulo 176 (CR, INB, MO), R. forest, páramo. Global distribution: Mexico to Brazil, Villalobos 288 (CR, INB). Greater Antilles, Azores. Records: A. Rojas 3236 (INB, MO), G. Davidse 28981 (MO).

Campyloneurum repens (Aubl.) C.Presl

Ceradenia kalbreyeri (Baker) L.E.Bishop forest, cloud forest, low elevation oak forest. Global Facultatively epiphytic herbs. Distribution in PILA distribution: Mexico to Brazil, Greater Antilles, Lesser 600–3100 m, low elevation mixed forest, mixed forest in Antilles, Africa, Madagascar. Records: A. Fernández 856 transition to low elevation forest, mixed forest in transition (CR, INB, MO).

to cloud forest, cloud forest, low elevation oak forest,

high elevation oak forest, páramo. Global distribution: Ctenitis equestris (Kunze) Ching

Guatemala, Costa Rica to Ecuador, Venezuela to French Terrestrial herbs. Distribution in PILA 1100–2000 m, Guyana. Records: G. Davidse 25249 (MO), G. Davidse mixed forest in transition to low elevation forest, mixed 28995 (MO), G. Herrera 5232 (CR, MO), G. Herrera forest in transition to cloud forest. Global distribution: 6222 (INB), J. Bittner 1427 (MO), J. Bittner 1766 (INB, Mexico, Honduras to El Salvador, Costa Rica to Colombia. MO), F. Quesada 1402 (INB). Records: D. Santamaría 3733 (INB).

Ceradenia phloiocharis L.E.Bishop Ctenitis hemsleyana (Baker ex Hemsl.) Copel. Epiphytic herbs. Distribution in PILA 2300–3200 m, low Terrestrial herbs. Distribution in PILA 1000–2700 m, elevation oak forest, páramo. Global distribution: Costa mixed forest in transition to low elevation forest, mixed Rica, Panama, Ecuador. Records: G. Davidse 25393 forest in transition to cloud forest, cloud forest, low (MO), G. Davidse 28979 (CR, MO), G. Davidse 28980 elevation oak forest, high elevation oak forest. Global (MO). distribution: Mexico to Panama. Records: A.K. Monro 5473 (BM, INB, MO, PMA), G. Herrera 5970 (CR, INB,

Ceradenia pilipes (Hook.) L.E.Bishop MO ), M. Moraga 497 (CR, INB). Epiphytic herbs. Distribution in PILA 2700–3100 m, low elevation oak forest, high elevation oak forest, páramo. Ctenitis melanosticta (Kunze) Copel. Global distribution: Costa Rica, Colombia to Bolivia. Terrestrial herbs. Distribution in PILA 200–1000 m, low Records: A. Rojas 3233 (INB), G. Davidse 25930 (MO), elevation mixed forest, mixed forest in transition to low G. Herrera 3428 (INB, MO). elevation forest. Global distribution: Mexico to Panama. Records: D. Santamaría 6145 (BM, INB, MO, PMA).

□ ε-PILA Ceradenia podocarpa (Maxon) L.E.Bishop Epiphytic herbs. Distribution in PILA 1600–2200 m, Didymochlaena truncatula (Sw.) J.Sm. mixed forest in transition to cloud forest, cloud forest, Terrestrial herbs. Distribution in PILA 0–2200 m, low low elevation oak forest. Global distribution: Costa Rica, elevation mixed forest, mixed forest in transition to low Panama. Records: G. Herrera 5381 (CR, INB, MO). elevation forest, mixed forest in transition to cloud forest, cloud forest, low elevation oak forest, pasture, dense

Ceradenia tristis A.R.Sm. thicket. Global distribution: Mexico to Argentina, Lesser Epiphytic herbs. Distribution in PILA 2800–2900 m, low Antilles, Greater Antilles, Asia, Africa, Madagascar. elevation oak forest, high elevation oak forest. Global Records: A.K. Monro 5699 (BM, INB, MO, PMA), A.K. distribution: Endemic to Costa Rica. Records: A. Rojas Monro 6166 (PMA), A. Soto 298 (INB), D. Santamaría 3239 (INB, MO). 5974 (BM, INB, MO, PMA), D. Santamaría 6159 (BM, INB, MO, PMA), D. Santamaría 6785 (BM, INB, MO,

Cochlidium rostratum (Hook.) Maxon ex C.Chr. PMA ), D. Santamaría 7225 (BM, INB, MO, PMA), D. Epiphytic herbs. Distribution in PILA 300–2700 m, low Santamaría 7452 (PMA), D. Solano 4284 (BM, INB, elevation mixed forest, mixed forest in transition to low MO, PMA), D. Solano 4311 (BM, INB, MO, PMA), D. elevation forest, mixed forest in transition to cloud forest, Solano 4887 (BM, INB, MO, PMA), G. Davidse 28446 cloud forest, low elevation oak forest, high elevation oak (CR, MO), G. Davidse 28512 (CR, MO), L. Acosta 2293 forest. Global distribution: Mexico to Venezuela, Greater (INB), R. Villalobos 279 (CR, INB, MO). Antilles, Lesser Antilles. Records: A. Fernández 1068 (CR, INB), G. Davidse 28781 (MO), G. Herrera 3424 Dryopteris flaccisquama A.Rojas (MO) , G. Herrera 5183 (CR, MO), G. Herrera 5391 (CR, Epiphytic herbs. Distribution in PILA 1500–2900 m, INB), G. Herrera 6162 (CR, INB, MO), H. Schmidt 649 mixed forest in transition to cloud forest, cloud forest, (INB), J. Bittner 1516 (MO), J. Bittner 1796 (INB, MO), low elevation oak forest, high elevation oak forest. Global J. Bittner 1797 (INB, MO), F. Quesada 1411 (INB), R. distribution: Mexico to Bolivia, Greater Antilles. Records: Aguilar 1166 (INB, MO). G. Davidse 28853A (MO), G. Davidse 29236 (CR, MO), G. Herrera 5493 (INB), M. Grayum 10289 (INB, MO).

Cochlidium serrulatum (Sw.) L.E.Bishop Facultatively epiphytic herbs.Distribution in PILA 0–2400 Dryopteris nubigena Maxon & C.V.Morton m, low elevation mixed forest, mixed forest in transition Terrestrial herbs. Distribution in PILA 3400–3500 m, to low elevation forest, mixed forest in transition to cloud páramo. Global distribution: Mexico to Panama. Records: