Liogenys foveata Cherman, 2021

Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes & Almeida, Lúcia Massutti De, 2021, Liogenys Guérin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) from the Chacoan Province and its boundaries: taxonomic overview with four new species, Zootaxa 4938 (1), pp. 1-59 : 19-21

Liogenys foveata Cherman , new species

Figs. 8 , 27 .

http://zoobank.org/ DB19CD93-CCCD-4A3F-A80E-74114513BDAC

Type material. Holotype labeled ( IADIZA): [white typeset] “ ARGENTINA: Formosa / RP 39, 23 km S Ing. Juárez, / 24º05ʹ27ʺS 61º56ʹ49ʺW. / 13-17-XII-08. F. Ocampo G. San Blas, F. Campón. ”, [red, typeset and handwritten] “ LIOGENYS FOVEATA / HOLOTYPE / Cherman M. A.”.

Holotype deposited at IADIZA.

Diagnosis. Body elongate; brown, elytra brown or light brown, pronotum purplish brown, head dark at vertex ( Fig. 8A ); clypeal emargination deep but not reaching the base of the teeth, angulate and wide; outer sides of anterior teeth parallel; clypeal lateral margin projection subangulate; outer side of anterior teeth and clypeal lateral projection forming an angle ( Fig. 8B ); fovea of the distal maxillary palpomere shallow, occupying almost the entire surface of the palpomere ( Fig. 8C ); pronotal posterior corners subangulate ( Fig. 8B ); mesofemural surface setose abundantly, long thick erect bristles, as well as the outer margin of the metafemur ( Fig. 8D, E ); pygidial width exceeding distance between spiracles of propygidium; pygidial disc with long bristles at the apex ( Fig. 8F ); parameres almost uniform along its length; outer margins slightly convex at the midline; apex spatulate ( Fig. 8G ).

Holotype. Male. Length: 10.5 mm; width: 5.6 mm. Brown. Head: distance between eyes almost twice the width of one eye; clypeal emargination deep but not reaching the base of the teeth, angulate and wide; outer sides of anterior teeth parallel; outer margin of anterior teeth almost equal to the eye length; lateral margin slightly convex, forming a subangulate projection; distance between clypeal lateral projection and anterior margin of eye shorter than the eye; distance between clypeal lateral projection and apex of the adjacent anterior tooth longer than the basal width of the anterior tooth; outer side of anterior teeth and clypeal lateral projection not forming an angle; distal maxillary palpomere, maximum width more than twice the apical width; fovea shallow, extending past the transverse midline, occupying almost the entire surface of the palpomere; antenna with 10 antennomeres, club lighter in color and slightly longer of the funicle. Thorax: anterior margin of pronotum straight; pronotal disc glabrous, punctures coarse and sparse; posterior corners subangulate; hypomere with short bristles, mesepisternum scaly; sides of metaventrite with bristles; distance between mesocoxae and metacoxae up to twice the length of the metacoxa; scutellum ogival, sparsely and finely punctate throughout. Elytra: shiny, glabrous, brown, lighter in color than pronotum; elytral suture darker than elytron and weakly elevated; all four elytral ridges weakly defined. Legs: three protibial teeth, the middle and apical equal in size; the three teeth equally spaced; mesofemural surface setose abundantly, as well as the outer margin of the metafemur, bristles long, thick, erect; mesotibia cylindrical in cross section, surface finely sculptured; two transverse carinae, the apical one present posteriorly; metacoxa with sparse, long scales; basal apophysis of metacoxa not produced; metatibia carinate along the inner margin towards the apex, apical inner surface setose; a metatibial transverse carina present posteriorly and posterior discontinuous longitudinal carina ( Fig. 8E ); metatibial apical spurs of different lengths; the larger equal in length to the diameter of the tibial apex; in males protarsomeres and mesotarsomeres I to IV enlarged; protarsomere II slightly longer than it is wide; metatarsomere I slightly shorter than metatarsomere II; claw bifid, symmetrical, superior tooth of a claw longer and slightly narrower than the inferior, distance between teeth equal to the length of the inferior tooth. Abdomen: ventrites with sparse, long bristles, abundantly on ventrite V; propygidium visible, with bristles; pygidium slightly convex, subquadrate, as wide as it is long; pygidial width exceeding distance between spiracles of propygidium; pygidial disc with long bristles at the apex; pygidial apex rounded. Parameres: basal region as wide as both sections of the parameres at its maximum width, parameral split at 2/3; inner margins straight, opened; outer margins slightly convex at the midline; apex spatulate; parameres slightly concave in lateral view ( Fig. 8H ).

Etymology. Adjective in the nominative singular. From the Latin foveatus (fovea “ditch, pit”): having a slight depression or pit in a bone or organ. In this species the fovea of the maxillary distal palpomere is shallow but quite broadly extended, almost reaching the apex.

Type locality. ARGENTINA, Formosa, Ruta Provincial 39, 23 km S Ingeniero Juárez, 24º05’27’’S 61º56’49’’W .

Geographical distribution. ARGENTINA (Formosa).

Remarks. Liogenys foveata resembles L. tarsalis , mainly in the size, body color, and clypeus deeply emarginate. Liogenys foveata differs from L. tarsalis (in parenthesis) mainly in the clypeus with emargination angulate and lateral margin with a weak subangulate projection (emargination subangulate, lateral margin with a distinct, rounded projection); in the large fovea on the maxillary distal palpomere, occupying almost the entire surface of the palpomere; in the pronotal posterior corners subangulate (rounded); and in the shape of the pygidium, tarsi and parameres. Females remain unknown.

Figures

FIGURE 8. Liogenys foveata Cherman, new species, male holotype views: A) dorsal, B) head and pronotum, C) distal maxillary palp, D) lateral, E) mesotibia and metatibia, F) pygidium, and parameres in views: G) dorsal, and H) lateral.

FIGURE 27. Geographical distribution of the Liogenys new species. Shaped points: new species; colors: Neotropical provinces (Morrone 2014) and Monte province from the South American Transition Zone (Roig-Juñent et al. 2018).