Magnacarina moderata Locht

Jorge I. Mendoza, Arturo Locht, Radan Kaderka, Francisco medina & Fernando Pérez-Miles, 2016, A new genus of Theraphosid spider from Mexico, with a particular palpal bulb structure (Araneae, Theraphosidae, Theraphosinae), European Journal of Taxonomy 232, pp. 1-28 : 21-25

publication ID	https://doi.org/ 10.5852/ejt.2016.232
publication LSID	lsid:zoobank.org:pub:52029B02-4A79-442D-8DA2-8DF3099ED8D0
DOI	https://doi.org/10.5281/zenodo.5631593
persistent identifier	https://treatment.plazi.org/id/74AF9502-86F8-4A92-B782-1A50555D8BD4
taxon LSID	lsid:zoobank.org:act:74AF9502-86F8-4A92-B782-1A50555D8BD4
treatment provided by	Jeremy
scientific name	Magnacarina moderata Locht
status	sp. nov

Treatment

Magnacarina moderata Locht View in CoL , Mendoza & Medina sp. nov.

urn:lsid:zoobank.org:act:74AF9502-86F8-4A92-B782-1 A50555 View Materials D8BD4

Figs 13–16 View Fig. 13 View Fig. 14 View Fig. 15 View Fig. 16

Diagnosis

Magnacarina moderata sp. nov. differs from all other Magnacarina species in lacking retrolateral nodule of megaspines and having only one prolateral accessory keel well developed. Magnacarina moderata sp. nov. also differs in possessing a secondary projection that is thinner and larger that of the other species, with a short and apically directed prolateral inferior keel. Females differ in carapace distinctly larger than wide.

Magnacarina moderata sp. nov. possesses the following character combination: male palpal bulb with a prominent primary projection strongly curved retrolaterally, with distal half slightly apically directed (Fig. A–C); retrolateral keel thin and slightly developed. Prolateral superior keel wide along its entire length, posterior extension ends almost in the middle of prolateral face ( Fig. 14 View Fig. 14 D–E); secondary projection is elongated and directed to apical, with one prolateral accessory keel well developed ( Fig. 14 View Fig. 14 C–D). Prolateral inferior keel wide, divided in two sections, increasing from back to front and distal end oriented to apical ( Fig. 14 View Fig. 14 A–B, D). Apical keel well developed, wider distally. Sperm pore is located between prolateral inferior and apical keels ( Fig. 14 View Fig. 14 D–E). Accessory apophysis slightly developed, bearing large megaspines at the apex ( Fig. 13 View Fig. 13 G–H). Metatarsus I lacking retrolateral nodule of megaspines ( Fig. 13 View Fig. 13 H). Spermatheca reduced, low, with one single receptacle oval in shape, strongly sclerotized. Uterus externus is three times wider, and four to five times longer than the spermatheca ( Fig. 15 View Fig. 15 E–F).

Etymology

The specific epithet is the latinized version of “moderate”, moderatus fem. moderata , referring to the shape of the secondary projection in the male palpal bulb which is moderately elongated and thinner than in other species.

Type material

Holotype

MEXICO: ♂, Nayarit, Acaponeta , 14 Nov. 1939, coll. C.M. Bogert and H.E. Vokes ( CNAN-T1003 ) .

Paratypes

MEXICO: 2 ♀♀, Sinaloa, 70 mi South of Mazatlan , 24 Jul. 1954, coll. W.J. Gertsch ( CNAN-T1004 and AMNH) .

Description

Holotype male CNAN-T1003 ( Figs 13– 14 View Fig. 13 View Fig. 14 )

Body length (not including chelicerae and spinnerets) 15.19, carapace: 7.31 long, 5.42 wide. Caput not markedly elevated; fovea procurved, 1.33 wide. Carapace with orange sheen setae ( Fig. 13 View Fig. 13 A). Eyes: anterior eye row procurved, posterior eye row, recurved. Eyes sizes and interocular distances: AME 0.24; ALE 0.32; PME 0.18; PLE 0.22; AME–AME 0.12; AME–ALE 0.08; PME–PME 0.46; PME–PLE 0.02; ALE–PLE 0.16. Ocular tubercle, width 1.18; length 0.84; clypeus 0.12 ( Fig. 13 View Fig. 13 D). Labium: length 0.90; width 1.175; with 58 cuspules. Maxilla inner corner with approximately 115-119 (left-right). Cheliceral promargin with 9-9 (left-right) teeth. Sternum length 3.07. Sigilla circular, second and third pairs hardly visible; posterior sigilla half its length from the margin ( Fig. 13 View Fig. 13 B). Leg formula: IV> I> III> II. Length of legs and palpal segments (femur, patella, tibia, metatarsus, tarsus, total): I: 5.29, 3.29, 3.41, 4.25, 2.26, 18.50; II: 4.71, 3.08, 3.33, 3.30, 2.64, 17.06; III: 4.18, 2.66, 3.03, 3.44, 2.55, 15.86; IV: 5.78, 3.13, 4.43, 4.95, 3.10, 21.39. Palp: 3.96, 2.82, 3.26, -, 1.29, 11.33. Spinnerets: PMS, 0.53 long, 0.20 apart; PLS, 0.97 basal, 0.63 middle, 1.10 distal. Tarsi I–IV scopulate, I and II divided by narrow band of setae, III and IV divided by strong band of setae. Metatarsi I scopulate on distal two thirds; II scopulate on distal half, III scopulate on distal half, and IV scopulate on distal fourth.

Tibia I with three apophyses, prolateral apophysis with one inner larger spine ( Fig. 13 View Fig. 13 G). Retrolateral apophysis slightly curved to dorsal ( Fig. 13 View Fig. 13 G, H). Accessory apophysis slightly developed, bearing four large megaspines at its apex ( Fig. 13 View Fig. 13 H). Metatarsus I curved ( Fig. 13 View Fig. 13 E), lacking retrolateral megaspines nodule ( Fig. 13 View Fig. 13 H). Retrolateral face of palpal tibiae with a short process, slightly developed on distal half ( Fig. 13 View Fig. 13 F). Chaetotaxy (left side): femora I 1 p; II 1 p; III 1 r; IV 2 r; palp none; patellae none; tibiae I 3 p, 4v; II 3 p, 8v; III 2 p, 7v, 3r; IV 9v, 4r; palp none; metatarsi: I 1 p, 2v; II 3 p, 8v; III 3 p, 8v, 2r; IV 2 p, 8v, 2r. Palpal bulb with a prominent primary projection strongly curved retrolaterally, with distal half slightly twisted to apical (Fig. A–C). Retrolateral keel thin and slightly developed. Prolateral superior keel wide along its entire length, posterior extension ends almost in the middle of prolateral face ( Fig. 14 View Fig. 14 D–E). Secondary projection elongated with one prolateral accessory keel well developed ( Fig. 14 View Fig. 14 B, D). Prolateral inferior keel wide, divided in two sections, increasing from back to front, distal end oriented to apical ( Fig. 14 View Fig. 14 A–B, D). Apical keel well developed, wider distally. Sperm pore is located between prolateral inferior and apical keels, and opens towards prolateral face ( Fig. 14 View Fig. 14 D–E). Tegular apophysis slightly developed ( Fig. 14 View Fig. 14 A–B). Urticating setae of type III arranged in one dorsomedian patch, with copper metallic iridescence ( Fig. 13 View Fig. 13 C). Variations summarized in Table 2 View Table 2 .

Color pattern: damaged by preservation in ethanol, carapace tegument looks yellowish; ventral coxae, labium, maxillae and sternum brown; abdomen dorsally brown with yellowish setae, ventrally brown. Urticating setae with copper metallic iridescence. Legs and palps: brownish.

Paratype female CNAN-T1004 ( Fig. 15 View Fig. 15 A–E)

Body length (not including chelicerae and spinnerets) 14.50, carapace: 6.85 long, 5.60 wide. Caput not markedly elevated; fovea procurved, 1.17 wide. Carapace with orange sheen setae ( Fig. 15 View Fig. 15 A). Eyes: anterior eye row procurved, posterior eye row, recurved. Eyes sizes and interocular distances:AME 0.225;

ALE 0.35; PME: 0.125; PLE: 0.275; AME–AME: 0.175; AME–ALE: 0.075; PME–PME: 0.55; PME–

PLE: 0.025; ALE–PLE: 0.10. Ocular tubercle, width 1.225; length 0.95; clypeus lacking ( Fig. 15 View Fig. 15 D). Labium: length 0.93; width 1.50; with 62 cuspules. Maxilla inner corner with approximately 134-123 (left-right). Cheliceral promargin with 9-9 (left-right) teeth. Sternum circular, length 3.10. Sigilla oval, second and third pair hardly visible; posterior sigilla once its length from the margin ( Fig. 15 View Fig. 15 B). Leg formula: IV> I> II> III. Length of legs and palpal segments (femur, patella, tibia, metatarsus, tarsus, total): I: 4.75, 3.17, 3.44, 2.53, 2.28, 16.17; II: 4.11, 2.93, 2.55, 2.39, 2.36, 14.34; III: 3.46, 2.43, 2.45, 2.90, 2.24, 13.48; IV: 4.99, 2.82, 3.85, 4.16, 3.05, 18.87. Palp: 3.43, 2.43, 2.29, -, 2.79, 10.94. Spinnerets: PMS, 0.87 long, 0.43 apart; PLS, 1.63 basal, 0.90 middle, 1.03 distal. Tarsi I–IV densely scopulate, I and II divided by narrow band of setae, III and IV divided by strong band of setae. Metatarsi I densely scopulate; II scopulate on distal two thirds, III scopulate on distal half, and IV scopulate on distal fourth. Chaetotaxy (left side): femora I 1 p; II 1 p; III 1 p, 1r; IV 1 r; palp 1p; patellae none; tibiae II 1v; III 2 p, 4v, 2r; IV 5v, 2r; palp 3v; metatarsi I 2v; II 4v; III 3 p, 7v, 2r; IV 3 p, 9v, 2r. Spermatheca fused, low with one single receptacle oval in shape, strongly sclerotized. Uterus externus is three times wider, and four to five times longer than the spermatheca ( Fig. 15 View Fig. 15 E–F). Urticating setae of type III, arranged in one dorsomedian patch, with copper metallic iridescence ( Fig. 15 View Fig. 15 C). Variations summarized in Table 2 View Table 2 .

Distribution

Known only from the border area between the states of Nayarit and Sinaloa ( Fig. 16 View Fig. 16 ), this species inhabits scrapes under rocks in deciduous forest.