Manota quantilla, Hippa, Heikki & Kurina, Olavi, 2013

Hippa, Heikki & Kurina, Olavi, 2013, New species and new records of Manota Williston (Diptera: Mycetophilidae) in the Neotropical region, Zootaxa 3686 (2), pp. 101-139 : 129-131

Manota quantilla sp. n.

Figs. 16 A–D

Male. Colour. Head dark brown, face brown. Antenna pale brown. Clypeus brownish, mouthparts pale yellowish. Thorax pale brown, medial part of scutum and scutellum at least seemingly darker brownish. Legs unicolorous yellowish. Wing pale brown; haltere yellow with dark brown knob. Abdominal tergites yellowish brown, sternites paler than tergites. All setosity pale, yellowish or brownish. Head. Antennal flagellomere 4, Fig. 16 A. Palpomere 3 of maxillary palpus with apicomesial thumb-like extension, with 3 apically curved sensilla; palpomere 4 with parasegment; palpomere 5 ca. 1.3 times longer than palpomere 4. Number of strong postocular setae 9. Thorax.

Anepisternum setose, with 32 setae; anterior basalare non-setose; preepisternum 2 non-setose; laterotergite nonsetose, episternum 3 setose, with 7 setae. Legs. Mid and hind tibial organs absent. Wing. R1 meeting C well on the basal half of the costal margin; the sclerotized part of M2 not extending to the level of the tip of R1; wing length 1.4 mm. Hypopygium. Figs. 16 B–D: Sternite 9 laterally fused with gonocoxa, the posterior margin transverse, slightly convex, extending to the middle of gonocoxa, anterior margin deeply incised, the setae few, similar to the adjacent ventral setae of gonocoxa. Ventral mesial margin of gonocoxa curved, the ventral setae of gonocoxa unmodified; posterolateral part of gonocoxa is drawn into a lobe. Parastylar lobe membranous or lacking, nonsetose. No paraapodemal lobe identifiable. Dorsal mesial margin of gonocoxa slightly lobe-like produced mesiad on the posterior half, with an aggregation of setae at the margin of the lobe, otherwise the setae on the dorsal side of gonostylus similar to those on the ventral side. Number of juxtagonostylar setae two, the ventral one flattened but otherwise rather unmodified megaseta, the dorsal one flattened with a whip-like branch, both arising from a common basal body which is about half of the length of the megasetae. Anteriorly from the juxtagonostylar megasetae there is a plate-like lobe bearing 1 large megaseta. Posteriorly from the juxtagonostylar megasetae there is a narrow transverse lobe bearing two or three strong curved setae. Gonostylus flat, bilobed, the smaller apicolateral lobe with two short setae, the larger apicomesial lobe with a few dorsal setae and with two long apical setae. Aedeagus broadly subtriangular, the lateral sides slightly concave, the apex curved ventrad. Hypoproct extending posteriorly to middle of gonostyli, the number of ventral setae (sternite 10) ca. 10 on each half. Cerci, except shortly separate apical parts, mesially fused.

Female. Unknown.

Discussion. By the non-setose anterior basalare, non-setose laterotergite, laterally with the gonocoxa fused sternite 9, membranous or lacking parastylar lobe, posterolateral lobe on the gonocoxa, subtriangular gonostylus with a long seta at the apicomesial angle and a curved seta at the apicolateral angle Manota quantilla resembles several species, viz. M. bisulca , M. costaricensis , M. depilis , M. fraterna , M. panda , M. papillosa , M. parva . It differs from all of them by having anteriorly from the juxtagonostylar megasetae, at the dorsal mesial margin of the gonocoxa, a lobe with only one large megaseta and without no other prominent setae or lobes. The row of 3 curved setae on the dorsal surface of the gonostylus is unique.

Etymology. The name is Latin, quantilla , how very little, referring to the small size of the fly.

Types. Holotype. Male, ECUADOR, Nape Prov., Yasuni NP, Yasuni Res. Station, 00°38’ S, 76°36’ W, Malaise trap, 3–20.xi.1998, leg. T. Pape & B. Viklund (on slide, in SMNH).

Figures

FIGURE 16. Manota quantilla sp. n. (holotype). A. Antennal flagellomere 4, latewral view. B. Hypopygium, dorsal view. C. Hypopygium, ventral view. D. Aedeagus and hypoproct, ventral view. Scale 0.1 mm.