Megazarinae Perrichot, 2009, 2024

Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo & Perrichot, Vincent, 2024, Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities, Arthropod Systematics & amp; Phylogeny 82, pp. 151-181 : 151

3.1. Subfamily † Megazarinae Perrichot, 2009 stat. nov.

Type genus.

† Megazar Perrichot, 2009.

Diagnosis.

Head hypognathous, globular; compound eye oval, not covering head length, without postocular carina; flagellomeres cylindrical, longer than wide; subantennal groove without dorsal carina; mandibles symmetrical, with three or four teeth. Posterolateral margin of pronotum with notch accommodating anterior thoracic spiracle, not surrounding spiracle posteriorly; mesoscutum large, around half length of mesosoma, parapsidal lines present; median mesoscutal sulcus and axillar grooves present, deeply crenulate or smooth. Metasoma elongate. Fore wing hyaline; C, Sc+R, M+Cu, A, Rs+M, cu-a, Cu, Rs and R1 pigmented; costal, radial, first cubital, submarginal, and marginal cells closed by tubular veins. Hind leg stout, metafemur and metatibia swollen; at least either inner margin of metatrochanter, metacoxa, metatibia, or metabasitarsus bearing row of comb-like, thick setae along ventral surface.

Genera included.

† Cretolyra gen. nov., † Genkyhag gen. nov., † Megacoxa gen. nov., † Megalava Perrichot, 2009 [† Megalavini trib. nov.]; † Megazar Perrichot, 2009 [† Megazarini Perrichot, 2009].

Stratigraphic extension.

Albian to Cenomanian.

Comments.

The tribe Megazarini Perrichot, 2009 is here elevated to subfamily level, as † Megazarinae stat. nov. based on the results of our phylogenetic analyses. The monophyly of the subfamily is supported by three synapomorphies under parsimony: (1) metatibia and/or metatarsus with comb-like setae along ventral surface (31:1; Fig. 3A ); (2) fore wing Rs+M located posteriorly, with the submarginal cell being at least twice the size of the medial cell (42:1; Fig. 3C ); and (3) fore wing pterostigma absent or reduced (46:0; Fig. 3C ). Within the subfamily, a new tribe is created to accommodate three of the newly described genera and † Megalava .

Figures

Figure 3. † Cretolyra noijebumensis gen. et sp. nov., holotype male IGR. BU- 067, mid-Cretaceous, Kachin amber. A habitus in lateral view; B head and mesosoma in dorsal view; C fore wing; D anterior part of mesosoma in lateral view with structures outlined; E metasoma and posterior part of the mesosoma in lateral view. Scale bars: 0.5 mm (A, B, C, E); 0.25 mm (D). Abbreviations: AG = inner axillar groove; MeS = mesometapectal sulcus; MMS = median mesoscutal sulcus; OC = occipital carina; PpL = parapsidal line; sp = anterior thoracic spiracle.