Minettia (Minettiella) membranacea, Li & Chen & Yang, 2020

Li, Wenliang, Chen, Xulong & Yang, Ding, 2020, Four new species of the subgenus Minettiella from China (Diptera, Lauxaniidae, Minettia), ZooKeys 932, pp. 93-111 : 93

Minettia (Minettiella) membranacea sp. nov. Figures 21-25 , 26-30

Type material.

Holotype: ♂ (CAUC), China, Yunnan: Baoshan Baihualing hot spring, 1500 m, 29.v.2007, Xingyue Liu.

Etymology.

Latin, membranacea , referring to the aedeagus with membranous processes internally.

Diagnosis.

Antenna brownish yellow, first flagellomere blackish brown on 2/3 apex of dorsal margin. Proboscis blackish brown, with pale yellow and black setulae. Mesonotum with two dorsocentral setae, acrostichal setulae in six irregular rows. Mid and hind tibia brownish yellow at base and tarsomeres dark yellow. Syntergosternite circular, broad dorsally and narrow ventrally, with one small anterior median process in lateral view, one membranous band under ventral bridge. Hypandrium median ventral side outward, V-shaped. Aedeagal apodeme long, slightly shorter than aedeagus.

Description.

Male. Body length 3.2 mm, wing length 4.3 mm. Female. Unknown.

Head (Fig. 21 ) black. Face and parafacial with dense grayish white pruinescence. Frons dark brown, as long as wide and parallel-sided, two wide black longitudinal stripes along fronto-orbital rows, extending from ocellar triangle to occiput; ocellar triangle black, ocellar setae developed, almost as long as posterior fronto-orbital setae and extending to anterior margin of frons; anterior fronto-orbital setae reclinate, shorter than posterior fronto-orbital setae. Gena ca. 1/7 height of eye. Antenna brownish yellow, first flagellomere blackish brown on 2/3 of the apex of dorsal margin, 1.7 times longer than high; arista blackish brown except paler basally, pubescent with longest setula almost as long as 1/3 height of first flagellomere. Proboscis blackish brown, with pale yellow and black setulae; palpus black with black setulae.

Thorax (Fig. 24 ) black, with brownish gray pruinescence. Two dorsocentral setae, acrostichal setulae in six irregular rows, with pair of strong acrostichal setulae before prescutellar setae; a pair of prescutellar setae, longer than anteriormost dorsocentral setae. One anepisternal seta and one katepisternal seta. Legs black, mid and hind tibia brownish yellow at base and tarsomeres dark yellow. Fore femur with eight posterior dorsal setae and six posterior ventral setae; fore tibia with one dorsal preapical seta and one short apical ventral seta. Mid femur with three anterior setae and one short apical posterior seta; mid tibia with one strong dorsal preapical seta and one strong apical ventral seta. Hind femur with one preapical anterior dorsal seta; hind tibia with one weak dorsal preapical seta and one short apical ventral seta. Wing slight yellow and hyaline, pale brown at base; costa with 2nd (between R1 and R2+3), 3rd (between R2+3 and R4+5), and 4th (between R4+5 and M1) sections in proportion of 9.6: 2.4: 1.8; r-m before middle of discal cell; ultimate and penultimate sections of M1 in proportion of 6.4: 4.4; ultimate section of CuA1 ca. 1/6 of penultimate. Haltere pale yellow.

Abdomen (Fig. 25 ) black, with sparse gray pruinescence. Male genitalia (Figs 26-30 ): syntergosternite circular, broad dorsally and narrow ventrally, with one small anterior median process in lateral view, one membranous band under ventral bridge. Epandrium slender. Surstylus separated from epandrium, with setae and curved in lateral view, with connective sclerite under cercus in posterior view; hypandrium median ventral side outward, V-shaped. Postgonite curved apically in ventral view, with long setae and nearly digitiform, aedeagus with membranous processes internally, with small apical concave; aedeagal apodeme long, slightly shorter than aedeagus.

Remarks.

The new species is very similar to Minettia (Minettiella) spinosa Shi & Yang from Hubei in the following characteristics: acrostichal setulae in six rows; the tarsomeres dark yellow, mid femur with three anterior setae and one short apical posterior seta; wing slight yellow and hyaline, but it can be separated from the latter in the face without spot; the mesonotum with pair of strong acrostichal setulae; the syntergosternite without setula near spiracle; the epandrium without subapical concavity; the hypandrium V-shaped. In M. (Minettiella) spinosa , the face with a yellow triangular median spot or only slightly yellow in center of face; the mesonotum without strong acrostichal setulae; the syntergosternite with one setula near spiracle; the epandrium with one small subapical concavity; the hypandrium H-shaped.

Distribution.

China (Yunnan).

Figures

Figures 21 - 25. Minettia (Minettiella) membranacea sp. nov. Male 21 head, anterior view 22 wing 23 habitus, lateral view 24 thorax, dorsal view 25 abdomen, dorsal view.

Figures 26 - 30. Minettia (Minettiella) membranacea sp. nov. Male 26 syntergosternite and epandrium, lateral view 27 epandrial complex, posterior view 28 syntergosternite, anterior view 29 aedeagal complex, ventral view 30 aedeagal complex, lateral view. Scale bar: 0.5 mm.