Mniotype dubiosa dubiosa (A. Bang-Haas, 1912)

Volynkin, Anton V., Matov, Alexei Yu., Behounek, Gottfried & Han, Hui-Lin, 2014, A review of the Palaearctic Mniotype adusta (Esper, 1790) species-group with description of a new species and six new subspecies (Lepidoptera: Noctuidae), Zootaxa 3796 (1), pp. 1-32 : 14-15

Mniotype dubiosa dubiosa (A. Bang-Haas, 1912) stat. n.

( Figs 43–45 , 81, 82 , 95 , 117, 118 )

Hadena lama var. dubiosa A. Bang-Haas, 1912 , Deutsche Entomologische Zeitschrift Iris 26: 149 (Type locality: „Mustagata Yarkend“ [NW China, Xinjiang, Keng Tau Mts, Muztagh ata Mt.]).

Type material examined. Lectotype. Designated herein for Hadena lama var. dubiosa : male, “Mustagata Yarkend” (printed) / “ Hadena Lama Stgr. v. Dubiosa BH” (handwriten) / “B.H.16” (handwriten) / “Preparation № MB.355 Ch. Boursin” / pink label “Origin.” (printed) / (the lectotype deposited in coll NKMB). Paralectotype: female, „Mus-tag-ata Yarkend“ (printed) / “ Hadena lama dubiosa ” (handwriten) / pink label “Origin.” (printed), slide Behounek 7659f (deposited in coll NKMB).

Other material examined. 1 male, 5.vii.2003, Kyrgyzstan, Sary-Dzhaz, floodplain of the river Kaingdy, near the mouth, V.V. Dubatolov, E.V. Nikolaeva [leg.] (Coll. SZMN); 1 male, 4.viii.2003, Kyrgyzstan, S shore of Issyk- Kul lake, E Kara-Beldek Mts over the Teploklyuchenka, upper support point of the forestry, at light, V.V. Dubatolov, E.V. Nikolaeva [leg.] (Coll. SZMN); 2 males, 19.vii.2003, Kyrgyzstan, Kashgaria, down stream of Archalsu river, V.V. Dubatolov & E.V. Nikolaeva leg. (Coll. SZMN); 1 male, [ Kyrgyzstan], 1906. vii.5, Semirech. area, Naryn. Akulin [leg.] (Coll. ZISP); 1 male, [ Kyrgyzstan], Semirech. obl., Naryn, vi.–viii.1905, unknown collector leg. (Coll. ZISP); 2 males, 4 females, [ Kyrgyzstan], Semirech. obl., Naryn, 10.v.-18.vi.1908, Akulin leg. (Coll. ZISP); 2 males, 4 females, 21-28.vi.1909 / Ulug-Tuz, Sary-Kol Ridge, East Turkestan, Divnogorsky [leg.] (Coll. ZISP); 4 males, 4 females, China, Xinjiang, SW from Kashi, Keng Tau Mts, Outag loc., N 38º54.363', E 075º13.788'', h- 2650 m, Floriani leg., 4–6.vi.2013 (Coll. ASV); 3 males, 1 female, Kazakhstan, Prov. Almaty, Zailiskiy Alatau, 1700 m, 15 km S of Issyk, Issyk Lake, N43º14', E077º28', 30.v.2006, leg. Balázs Benedek (Coll. BBT); 1 male, 07.vii.2011, E Kazakhstan, Zaisan distr., Saur Mts, 24 km S Zaisan, upper stream of Bolshoi Zhemenei river, 1650 m, 47°14’ N, 84°56’ E. Titov S.V. leg. (Coll. AVB); 2 males, same locality, 17–19.vii.2011, Yakovlev R.V. leg. (Coll. AVB); 3 males, 1 female, 18–19.vii.2011, same locality, Titov S.V. leg. (Coll. AVB); 1 male, 09.vi.2013, E Kazakhstan, East Kazakhstan area, Urdzhar district, Tarbagatai Ridge, 6,7 km N of Kyzymbet (Alekseevka) village, mesophilous shrubby slopes, 1300 m. 47°18.365’ N, 81°32.152’ E, Volynkin A.V., Titov S.V. & Černila M. leg. (Coll. AVB); 1 male, 15.viii.2010, SE Kazakhstan, Zailijsky Alatau Ridge, middle stream of Assy river, 1700 m, Zhaman-Uj gorge, Egorov P.V. leg. (Coll. AVB); 1 female, 22–23.vii.2009, SE Kazakhstan, Ketmen' Ridge, Amirsaj, Taranov B.T. leg. (Coll. AVB).

Slides AV0680, AV0685, AV0693, AV0694, AV0695, AV0704, AV0714, AV0715, AV0807, AV1123 Volynkin (males), AV0716, AV0878 Volynkin, OP2299 f, OP2300 f Pekarsky (females).

Note. Previous authors have treated dubiosa as a subspecies ( Hacker & Peks 1990; Ronkay et al. 2001) or a synonym (Hacker 1990) of M. lama . Study of the type material demonstrates its specific status.

Diagnosis. Wingspan 37–40 mm. Head, thorax and abdomen brown. Ground colour of forewing brown. Outer margin of reniform dentate. Medial area shadowed around claviform and medial fascia. Terminal area dark brown. Hindwing pale brown. This species is externally close to M. juldussica , but differs in the less dentate submarginal line, darker and more monotonous colouration of forewing and less expressed pattern. The correct determination of pale or damaged specimens often requires the study of the genitalia. The male genitalia of M. dubiosa ( Figs 81, 82 , 95 ) differ clearly from those of M. juldussica in the narrower juxta with shorter serrated parts of its apical processes (in M. juldussica distal half, but in M. dubiosa only distal third serrate), narrower clavus strongly protruding from surface of sacculus, shorter ampulla, more rounded and apically elongated cucullus, obtuse and strongly dentate apical part of carina, shorter and less sclerotised eversible dorso-lateral bar of carina arising from the apical part of carina, and larger band of spinules on terminal diverticulum. The female genitalia ( Figs 117, 118 ) differ in the narrower ostium bursae, narrower and less curved ductus bursae, weakly sclerotised appendix bursae and posterior part of corpus bursae, larger conical anterior projection of anterior part of corpus bursae and smaller signum.

Distribution. Central Asian. East (Saur and Tarbagatai Mts) and South-East Kazakhstan, Tien Shan massif, Pamir and Alai Mts, nortwestern China (Xinjiang).

Figures

FIGURES 41 – 50. Mniotype spp. adults: 41, M. anilis anilis, male, Italy, Lazio region (GBG / ZSM); 42, M. anilis sylvatica, male, France, Corsica (GBG / ZSM); 43, M. dubiosa dubiosa, lectotype male, " Mustagata Yarkend " (NKMB); 44, ditto, male, Tajikistan, Pamir Mts (ZISP); 45, ditto, female, SE Kazakhstan (AVB); 46, M. dubiosa amitayus, holotype male, China, Tibet Plateau (PGM); 47, ditto, paratype female, China, Sichuan Prov. (BBT); 48, ditto, paratype male, Bhutan (URK); 49, M. krisztina, male, China, Tibet Plateau (BBT) (Photo by B. Benedek); 50, ditto, female, China, Tibet Plateau (CAU).

FIGURES 81 – 86. Mniotype spp., male genitalia: 81, M. dubiosa dubiosa, lectotype, Muztagh-Ata Mt., slide MB 355 Boursin; 82, M. d. dubiosa, Kyrgyzstan, slide AV 0693 Volynkin; 83, M. d. amitayus, holotype, China, Tibet Plateau, slide 3210 Gyulai; 84, M. d. amitayus, paratype, Bhutan, Prov. Paro, slide AV 1116 Volynkin; 85, M. krisztina, W China, slide hhl- 2162 - 1 Han; 86, M. krisztina, W China, slide hhl- 2272 - 1 Han.

FIGURES 87 – 96. Mniotype spp., male genitalia: carina. 87, M. lama lama, Kyrgyzstan, slide AV 0681 Volynkin; 88, M. adusta adusta, E Kazakhstan, slide AV 0713 Volynkin; 89, M. adjuncta adjuncta, Pakistan, slide AV 0792 Volynkin; 90, M. juldussica, Tajikistan, slide AV 0796 Volynkin; 91, M. kobyakovi, paratype, Altai Mts, slide AV 0744 Volynkin; 92, M. bathensis, Russia, Altai Mts, slide AV 0513 Volynkin; 93, M. aulombardi, Taiwan, slide AV 0786 Volynkin; 94, M. anilis anilis, Italy, Reg. Lazio, slide GB 8141 m Behounek; 95, M. dubiosa dubiosa, E Kazakhstan, slide AV 0680 Volynkin; 96, M. krisztina, China, Tibet, slide hhl- 2506 - 1 Han.

FIGURES 113 – 120. Mniotype spp., female genitalia: 113, M. bathensis, South Siberia, slide AV 0725 Volynkin; 114, M. aulombardi, paratype, Taiwan, slide 8144 f Behounek; 115, M. anilis anilis, France, slide GB 8142 f Behounek; 116, M. a. sylvatica, Corsica, slide AV 0787 Volynkin; 117, M. dubiosa dubiosa, paralectotype, " Mustagata Yarkend ", slide 7659; 118, ditto, SE Kazakhstan, slide AV 0878 Volynkin; 119, M. d. amitayus, paratype, W China, Tibet Plateau, slide AV 0777 Volynkin; 120, M. krisztina, China, Tibet, slide hhl- 2271 - 2 Han.