Mycomya naja, Väisänen, Rauno, 2013

Väisänen, Rauno, 2013, New Mycomya species from the Himalayas (Diptera, Mycetophilidae): 3. Subgenera Cesamya and Mycomyopsis, Zootaxa 3737 (2), pp. 129-153 : 146-148

Mycomya naja sp. n.

Figs. 12 A–G

Material studied. Holotype. ♂. NEPAL, Ktmd., Pulchauki, 6800', Mal. tr., 14.VII. 1967, Can. Nepal. Exp. (in CNC). Paratypes (only ♂♂). Same data as holotype, but 6600', 10.VIII. 1967, 1 ♂ (in CNC); Ktmd., Godavari, R. Bot. Gdn., 5000', 29.VII. 1967, Can. Nepal Exp., 1 ♂ (in MZH); Dobang Kharka, 83°24'E, 28°36'N, 2400 m, 9.X. 1971, A. Nakanishi, 2 ♂♂ (in KUC); same data, but 10.X. 1971, 1 ♂ (in KUC); same data, but 15.X. 1971, 1 ♂ (in KUC); same data, but 20.X. 1971, 1 ♂ (in KUC); same data, but 28.X. 1971 (1 ♂ KUC); same data, but Malaise trap, 25–26. X. 1971, 1 ♂ (in KUC); Bogara, 83°23'E, 28°35'N, 1700–2200 m, Malaise trap, 18–19. IX. 1971, A. Nakanishi, 1 ♂, 1 ♀ (in KUC); same data, but 19–20. IX. 1971, 4 ♂♂ (in KUC); same data, but 20–21. IX. 1971, 5 ♂♂ (in KUC), same data, but 21–22. IX. 1971, 2 ♂♂ (in KUC); Lipshiba Kharka, 83°23'E, 28°35'N, 30.X. 1971, A. Nakanishi, 1♂ (in KUC); Basantapur, 27°06'N, 87°23'E, 2300 m – 27°08'N, 87°26'E (specimen collected on the way between the two localities), 26.VI. 1972, H. Shima, 1 ♂ (in KUC); Thurukpa, 27°36'N, 87°36'E, 2600 m – Papun, 27°32'N, 87°38'E, 2100 m (specimen collected on the way between the two localities), 10.VII. 1972, J. Emoto, 1 ♂ (in KUC); Topke Gola, 27°38'N, 87°35'E, 3700m – Thurukpa, 27°36'N, 87°36'E, 2600 m (specimen collected on the way between the two localities), 9.VII. 1972, P. Norbu, 1 ♂ (in KUC); Gupa Pokali, 27°17'N, 87°30'E, 2900 m – Chouki, 27°12'N, 87°28'E, 2700 m (specimens collected on the way between the two localities), 16.VII. 1972, J. Emoto, 2 ♂♂ (in KUC). Other material. MYANMAR, Kambaiti, 2000 m, 31.III. 1934, Malaise, 1 ♂ (in MZH); same data, but 4.VI. 1934, 1 ♂ (in MZH).

Description. ♂. Head. Palp brownish, other mouthparts yellowish, face yellow, posterior parts of head brownish. Antenna brownish, scape, pedicel and base of 1st flagellomere yellow. Length of 1st flagellomere about 3.5x, 2nd flagellomere 2x, its width. Thorax. Pronotum yellow, with 2 long setae. Scutum yellow to yellowish with 3 brown longitudinal, posteriorly fused stripes. Anepisternum yellowish. Preepisternum yellow, ventral part light brownish. Scutellum yellowish, with 4 long setae. Laterotergite brownish. Mediotergite brownish, bare. Wing. Length 3.3–5.0 mm. Wing hyaline. Sc ending in R1 near or proximad of middle of small cell, Sc1 missing. Apical part of Sc bearing 0–1 small setae. Small cell 1–1.5x as long as wide. Cu fork distinctly distal to M fork. M ratios: 1.00–1.09, 1.38–1.50. Cu ratios: 0.93–1.14, 1.56–2.08. Small setae: M petiole: 0; M1: 0–7; M2: 0–4; Cu petiole: 0– 1; Cu1: 0–12; Cu2: 0–7. Halter pale yellowish. Legs. Coxae yellow, femora yellow, tibiae and tarsi brownish to brown. Coxa 2 without spur. Leg ratios: bt1:t1 = 0.82–0.87, bt2:t2 = 0.65–0.73, bt3:t3 = 0.58–0.60. Abdomen. Tergites brownish, sternites brownish with yellowish posterior margins. Hypopygium. Figs. 12 A–G: yellowish. Tergal part with 2 pairs of combs, outer combs very narrow and their base deeply bilobed ( Fig. 12 A). Tergal lateral appendage setose, wide, basally more than 2x as wide as its slender apical part; apical part of tergal lateral appendage narrow, strongly curved, with some flattened, curved apical setae ( Fig. 12 D). Sternal submedian filament relatively short, slightly curved, extending slightly beyond apex of sternal synsclerite. Gonostylus more than 4x as long as wide, apically slightly wider than basally, with 3–4 subapical teeth and long, strong, slender spur with 1 apical tooth, and membranous lateral lobe about ½ of length of main branch of gonotylus ( Fig.12 E). Aedeagus with 2 lateral appendages extending slightly beyond its apex ( Fig. 12 F). Female. Wing length 4.7 mm. Thorax yellow, posterior part of mediotergite brownish. Abdomen brownish. Veins Sc, M and Cu bare.

Discussion. The male hypopygium of M. naja is generally quite similar to Chinese M. vaisaneni Wu & Yang, 1994, but differs from it by having a strong spur on the gonostylus ( Fig. 12 E), similar to that of the Nearctic M. sublittoralis Shaw, 1941 . Mycomya sublittoralis is otherwise very different from M. naja , e.g. its tergal combs are much wider than in M. naja ( Fig. 12 A) and the setosity of the tergal lateral appendage is not restricted only to the basal and apical parts (Väisänen 1984: figs. 1051–1056).

Etymology. The species name is a noun in apposition and refers to Naja Laurenti, the genus for the cobras, a group of venomous elapid snakes.

Figures

FIGURE 12. Mycomya naja sp. n., male hypopygium. A – C tergal, sternal and lateral views, D tergal lateral appendage, twice enlarged, E gonostylus, twice enlarged, F apex of aedeagus, sternal view, twice enlarged, G. tergite 8. Scale 0.2 mm.