Naarda laoana, Tóth & Ronkay, 2015

Tóth, Balázs & Ronkay, László, 2015, Revision Of The Palaearctic And Oriental Species Of The Genus Naarda (Lepidoptera: Erebidae, Hypeninae) Part 2. Description Of Ten New Species From Asia, Acta Zoologica Academiae Scientiarum Hungaricae 61 (1), pp. 3-23 : 5-9

Naarda laoana sp. n.

( Figs 2 , 14 , 26, 29)

Holotype. m, Laos: Prov. XiangKhouang, PhouSamson , 10 kmEBanPeun, 19°8.717’N, 103°46.811’E, 15.II.2012, Lao 2012PL_20, leg. L. Peregovits & A. Szappanos; slideNo. TB 555m (coll. HNHM) .

Paratypes. 4 m, 1 f, dataasholotypebutslideNo. TB 554m, TB556f (coll. LP) .

Taxonomy. Thisspeciesistheclosestrelativeof N. octogesima .

Description. Wingspan 22–25.5 mm, lengthofforewing 11–12.5 mm. Oneofthelargest Naarda species. Antennaefiliform, ciliate, setosewithonesetapersegment. Lengthof setae 1.5–2 timesdiameterofflagelluminmale, nearlyorjustaslongasdiameterofflagelluminfemale. Ciliaofmalesaslongasorsomewhat (ca 1.5 times) longerthandiameter offlagellum, arrangedtofourtransversearrays; ciliaoffemaleshalfaslongasdiameter offlagellum, inthreetransversearrays. Maleantennawithlongscalesformingasmall nodeonthecentralsegments. Labialpalpsrelativelyshort, theirlength 2.5 timesdiameter ofeyeinmale, 3.5 timesinfemale. Palpsbroadestatapicalpartinmale, atcentralpartin female; dorsaledgeofthelatterbeingstronglyconvex. Scale-hoodofvertexwide, itstip rounded, longinmale, shortinfemale (halfaslongasinmale). Protibiaeinmaleslightly broader, hairy. Characteristicwingpatternfeatures: forewingcostanotconcaveinmale; groundcolourochreousbrowninmale, darkchestnut- browninfemale, basalareasuffusedwithyellowishinbothsexes; subterminallinegreyishwithtwoprojectionsinwards inbothsexes, itscolourlighterinfemale, postmediallinestrictlyparallelwithsubterminal inbothsexes, angledinwardsandalmosttouchingreniformstigmainmale; medialline dark, wide, slightlysinuousinmale, straightinfemale; antemediallinepresent. Reniform stigmabig, angularandoblong, darkochreousyellow, usuallyobsolescent, lessprominent despiteitssize; withtwodarker, brown-shadeddotsinupperandlowerthirds, bottom onesometimesdarkerthantopone; orbicularstigmaquitebig, asyellowasreniform, with broad, darkoutline. Hindwinggroundcolourslightlymoregreythanthatofforewing, discalspotabsent, subterminallinemuchlighterthangroundcolour.

Malegenitalia ( Fig. 2 ). Uncusrelativelylong, curved, withabulgeattheventraledge ofthebasalhalf, itstippointed. Scaphiumaslongasuncus, butcurvedagainstit. Tegumenlongerthanvinculum; thelatterstronglysclerotised, withslenderventralprocess. Saccusbroad-based, abruptlytapering, long, itsdistalhalfquitenarrow, itstiprounded. Baseoftranstillaverybroad. Juxtatriangularorheart- shaped (taperingventrally), with smallventro- medialnotch. Valvabroad-based, abruptlytaperingtowardssubapicalneck. Cucullusverynarrow, freeonlyattip. Sacculuswelldeveloped, distallydilated, itsdistal endtwicebroaderthanbase. Harpearelativelybroad-basedprocess, itstiprounded, its basefusedwiththeventro- lateralcornerofthesacculus. Costaheavilysclerotised, itsdorsaledgestraight, distalendbifurcate, moreorlessbird-head-like, withlong, acuteventral extension (“bill”) whichdoesnotsurpassingventraledgeofcucullus; dorsalextension rounded, tongue-shaped. Aedeagusstraight, verythick; vesicasphaericalwithtwosmall diverticula, itssurfacesmooth, nearlytwicebroaderandlongerthanaedeagus, withan extremelylarge, broadandstraight, apicallybifurcatecornutus.

Femalegenitalia ( Fig. 14 ). Ovipositorlobesangular. Apophysesposterioresslightly (ca 1.3 times) longerthanapophysesanteriores. LobesofsternumA7 rounded. Sinusvery broad. Ductusbursaeveryshortandsurprisinglynarrow, hardlysclerotised. Corpusbursaeoval, itsanteriorquarterdenselyscobinatebyverysmallgranules. Astronglysclerotised, broad, longitudinalbandpresentfromthemouthofductusbursaetoanteriortipof corpusbursae, endinginasmallappendix.

Diagnosis. Thecomparisonofthisnewspecieswithitsclosestrelative, N. octogesima , isgivenundertheDiagnosisofthatspecies. Thebestdistinctiveexternalfeaturesof N. laoana arethelessprominentreniformstigma, the moreyellowishsuffusedbasalpartofbothwings, andthemoredistinctly markedyellowsubterminallines. Thekeyfeaturesofthemalegenitaliaare thestraightandapicallybifurcatecornutus, thelackofscobinationofvesica, thelargerandmorebird-head-likesclerotisedendofcostawithlongerand moreacuteventralextensionandthefiner, thinnervinculum. Thefemale genitaliaof N. laoana areeasilyseparablefromthoseof N. octogesima bythe shorterandthinner, membranousductusbursaewithshorteranteriorpocketlikeextension, andthelessscobinatecorpusbursae.

Themalegenitaliaof N. laoana resemblethoseof N. abnormalis buthave narrower, longeruncusandscaphium, muchlongersaccus, muchmoredevelopedcosta, muchbroaderdistalpartofsacculusandsmallerharpe. The twospeciesaremoredissimilarexternally, thenewspeciesismuchlargerin size, andthecostaofmaleisnotconcaveincontrasttothelatterspecies. The groundcolourof N. laoana islighter (inmale) ormorereddish (infemale), the reniformstigmaislargerbutlessconspicuousthanin N. abnormalis .

Etymology. Thisisthemostinterestingspeciesamongthefew Naarda taxadescribed fromLaos.

Distribution. Laos, southernpartoftheXiangkhoangPlateau.

Figures

Figs 1–6. Male genitalia of newly described Naarda species: 1 = N. octogesima sp. n. (RL7903m), 2 = N. laoana sp. n. (TB554m), 3 = N. mirabilis sp. n. (RL7920m), 4 = N. vicina sp. n. (clasping app.: RL7988m; aedeagus: RL7989m), 5 = N. atrata sp. n. (TB392m), 6 = N. furcatella sp. n. (clasping app.: TB447m; aedeagus: TB419m).

Figs 13–18. Female genitalia of newly described species: 13 = N. octogesima sp. n. (TB399f), 14 = N. laoana sp. n. (TB556f), 15 = N. vicina sp. n. (RL8000f), 16 = N. atrata sp. n. (TB448f), 17 = N. plumbea sp. n. (RL7917f), 18 = N. furcatella sp. n. (TB422f).