Nanhaia speciosa (Champ. ex Benth.) J.Compton & Schrire, 2019

Compton, James A., Schrire, Brian D., Koenyves 3, Kalman, Forest, Felix, Malakasi, Panagiota, Sawai Mattapha, & Sirichamorn, Yotsawate, 2019, The Callerya Group redefined and Tribe Wisterieae (Fabaceae) emended based on morphology and data from nuclear and chloroplast DNA sequences, PhytoKeys 125, pp. 1-112 : 38-39

Nanhaia speciosa (Champ. ex Benth.) J.Compton & Schrire comb. nov.

≡ Millettia speciosa Champ. ex Benth., Hooker’s J. Bot. Kew Gard. Misc. 4: 73 (1852). Type: China, "Hong Kong, Millett 505", K000881029 (K, lecto.!, designated by Schot (1994) but see note below)

≡ Phaseoloides [ Phaseolodes ] speciosa (Champ. ex. Benth.) Kuntze, Revis. Gen. Pl. 1: 201 (1891)

≡ Callerya speciosa (Champ. ex Benth.) Schot, Blumea 39(1-2): 32 (1994)

Nomenclatural note.

Schot selected a specimen at K, Champion 505 as lectotype for the name Millettia speciosa Champ. ex Benth. ( Schot 1994: 32). This was, however, clearly an error because another specimen, Millett 505, was cited in the protologue and determined as type by Dunn’s annotation on the sheet at K prior to the publication of his monograph on Millettia in 1912. Dunn had incorrectly cited the specimen as Champion 505 and not Millett 505 which may explain Schot’s error ( Dunn 1912a: 155). Bentham’s paper on the plants of Hong Kong was based on the collections of Major John George Champion (1815-1854). There are two specimens at K collected by Champion in the Hooker Herbarium: K000881027 and K000881028. Both are annotated " Champion 261, Hong Kong", but these specimens were not specifically cited by Bentham in the protologue which merely stated "on Victoria Peak" [on Hong Kong island] and are therefore regarded as uncited specimens that comprise part of the original material associated with the name (Art. 9.4(a)). The Millett collection K000881029 is annotated "505 Millettia grandiflora sp. n. fls white and yellow Hong Kong" and is dated 1854 in Bentham’s Herbarium. This collection was cited in the pro tologue "it is in the Hookerian Herbarium from Millett’s collection" and is therefore a syntype in the absence of any cited holotype (Art. 9.6). Under Art. 9.12 a syntype takes precendence over any uncited specimens when selecting a lectotype ( Turland et al. 2018). Millett 505 is therefore the obligate lectotype (N. Turland pers. comm.).

Illustrations.

Lôc and Vidal in Fl. Cambodge, Laos & Vietnam 30: 37 (2001); Wei and Pedley Fl. China 10: 182, t. 215 [1-7] (2010). http://www.fpcn.net/a/tengmanzhiwu/20131008/Callerya_speciosa.html (as Callerya speciosa ). Fig. 4 .

Distribution.

China (Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hunan, Yunnan); Vietnam (north).

Habitat.

In open forest, edges of ravines and thickets from 100-300 m.

Figures

Figure 4. Nanhaia speciosa (Champ. ex Benth.) J. Compton & Schrire. A Habit B flower bud with bract and bracteole C calyx exterior and bracteole D detail of calyx exterior E detail of calyx interior F standard petal G wing petal H keel petal I staminal column J staminal column lateral view K stamen ventral and dorsal view L ovary and style M style and stigma N pod O seed lateral view P seed ventral view (all from Shiu Ling Hu 6091). Drawn by Margaret Tebbs.