Eunatalis spinicornis (Blackburn, 1890)

Gerstmeier, Roland & Seitner, Martin, 2013, Revision of the checkered beetle genus Eunatalis Schenkling, 1909 (Coleoptera: Cleridae: Clerinae), Zootaxa 3698 (1), pp. 1-77 : 63-66

Eunatalis spinicornis (Blackburn, 1890)

( Figures 49–50 , 58 B)

Natalis spinicornis Blackburn, 1890: 123 ; Blackburn 1899: 29.

Specimens examined: Lectotype (designated here): Australia., Blackburn Coll., B.M. 1910-236, Natalis spinicornis Blackb. ; Type (BMNH). Paralectotype (designated here): Adelaide, Blackburn; Natalis spinicornis co-type; Natalis spinicornis Bl, S. Australia, Cotype; No. 25-035814 (SAMA).

Comment on the type specimens: Although the BMNH specimen is marked with Blackburn's distinctive "T" (indicating the holotype) it must be, along with the SAMA specimen, considered a syntype because Blackburn (1890: 123) failed to designate type status to either specimen within in his published description. Both specimens are available for lectotype designation (ICZN 1999: Article 74.1).

Others: Lake King (Strandline), West Aust., 8. Feb. 2000, P. Hutchinson; Reg.no. 833303 (WAMP). WA: State Library, Northbridge (Perth), 31°56'56"S, 115°51'37"E, 22 March 2011, via S. Crockett, Reg.no. 83313 (WAMP). M.L. July 35, Evansford; Brit. Mus. 1947-431; E. spinicornis, J.J. Menier '88 (BMNH). Natalis porcata F., Gawler (NMPC). Broken Hill., 16.3.(?), C.E. Chadwick; Eunatalis porcata Fabr., Det. by K.C. McKeown (ANIC). Australia., H.J. Nillier., 1905-232; Killalpanima., 100 miles E. of L. Eyre; Natalis spinicornis Blackb., S. Schenkling det. (BMNH). Natalis psp., Id. by R. Harvey; 201; Ex. Coll. R.V. Southcott; No. 25-036016; Glenunga, 26.8.37 (SAMA). Rockhampt., Heyne Broth.; Coll. S. Schenkling; Schenkling det; Col-02898, 3 ex. (SDEI). Rockhampton (ZMB), (MNHN). Rockhampton, Queensland; 25-035878 (SAMA). VIC: Koorlong, Plot B from Trap Logs, 21 Feb 1995, Emerged 26 Jul 1995; Eunatalis spinicornis Blackburn , det. T.A. Weir 1996 (ANIC). Adelaide, Aug. 1956; C.H.S. Watts; Natalis spinicornis B., id. by C. Watts; No 25-035892 (SAMA). Adelaide, Fren (?); Schenkling det; Eunatalis spinicornis (Blackb) , Ginter Ekis det. 1985 (SDEI). Adelaide, SA 1906; Coll. Hacker; Schenkling det; Col-02896 (SDEI). Adelaide; Natalis spinicornis Blackb. (BMNH). Adelaide, July 1955; No. 25-036034 (SAMA). Adelaide; No. 0 35894 (SAMA). Adelaide, June 1955; C.H.S. Watts; No. 25-035889 (SAMA). Adelaide, A.M. Lea; No. 25-035883 (SAMA). Murray. R., S.A., E.R. Zietz; A.H. Elston Collection; K 324408; Natalis spinicornis Bl., Id. by A.H. Elston (AM). Australia: W.A., New Norcia, IV-18-1964, H. Demarz leg; 8 ex. (FMNH). Corrigin, Valema Farms, W.A., P#3, active 14 Oct. 2001, caught in net Golding/Pitman; Reg.no. 83306, 83307, 83308; 3 ex. (WAMP). Murrumbateman, NSW, E.C. Zimmermann, 29-IV-89; Brit. Mus. 1989-119 (BMNH). Kellerberrin, W.A., H.M. Gilles, 3.2.0 7 and 6.2.07; G. Bryant Coll. 1919-147; 2 ex. (BMNH). Kellerberrin, W.A., H.M. Gilles, 4.2.07; Natalis spinicornis Blackb. (BMNH). Kellerberrin, W.A., H.M. Gilles, 1.2.07; Natalis spinicornis Blackb. ; 2 ex. (BMNH). Black Mt. A.C.T., light trap, 2.2., 19.2., 22.2.,4.3., 31.3.68, M.S. Upton; 5 ex. (ANIC). Black Mtn. ACT, 28 Dec. 1967, K. Pullen (ANIC). Mt. Lofty Rgs., S.H. Curnow; A.H. Elston Collecton; K 324404 (AM). Mount Lofty; S.H. Curnow Collection; No. 25-035951 (SAMA). Mt. Lofty Rgs., S.H. Curnow; No. 25-035982 (SAMA). Mt. Gambier; Mus. Paris, Coll. Castelnau, Coll. Sedillot 1935 (MNHN). WA: Kalbarri Caravan Park, 27°41'58"S, 114°10'16"E, 5 March 2004 (elev. 20m), G. Byrne '455; 5:30 pm, fine, hot windy, under gas stove, in dark; Reg.no. 83311 (WAMP). S. Australia: Robertstown, Brady Creek, iv.1936, T. Honeychurch (BMNH). Marloo Stn,, Wurarga, W.A., 1931–1941, A. Goerling; 2 ex. (ANIC). S. Aust., 36 km ESE Amata, Malaise Trap 21–24 Oct. 98, 26°16'12"S 131°28'36"E, Pitjantjatjara Land Surv YUROS; E. camaldulensis, Melaleuca glomerata creekline, No. 25-035901 (SAMA). S. Aust., Tarlee, 17 Feb. 1974, for P.B.McQuillan; No. 25-035916 (SAMA). N.H. Swan R. (BMNH). Hermannsburg, N.T., S. Australia, H.J. Nillier, 1907-283 (BMNH). Melbourne, D str ct V, R. Blackwood; K 324380, 324367; 2 ex. (AM). Melb.; spinicornis Bl; Col-65749 (NMV). Rockingham: Titana Thoms. (ZMB). Barakula, april 1939; Eunatalis spinicornis , det. J.S. Bartlett 15. Aug. 2008; QDPC O-166562 (QDPC). Winston Hills, NSW, 16.iii.1996, C.A.P. Vranhurst, K 324483 (AM). Dunmore, via Dalby QLD., 4.iii.1980, 20.iii.1980, M. De Baar S. Fatt; Ex Malaise Trap, Accn.No. 1818-33; QDPC O-166583; 2 ex. (QDPC). Natalis fasciata, Kunwarara , 6-5-35; S.R.E. Brock Collection donated to ANIC 1987; 2 ex. (ANIC). W.- Australien, Coolgardie, 5.II.67, leg. H. Demarz (ZSM). Coolgardie; No. 25-035851 (SAMA). Dawson distr., (Barnard coll.) (MNHN). Launching Place V., C. Oke, Col-65748 (NMV). Bendigo, Vic, C. Oke; Col-65749 (NMV). Heyfield 23.3.18; Col-65755 (NMV). Alice Springs, N.T., Demarz, 8.1965; Australien, leg. Demarz (NHMB). near Alice Springs, M.W. Mules; No. 25-035928 (SAMA). New South Wales, Barham, January 1969, Coll. Rudolf Komout (ANIC). 35°16'57S 149°06'22E, Black Mountain, CSIRO, site ACT, 4 Feb 1998, M. Neave & S. Grosse; 2 ex. (ANIC). 31.11S 121.38E WA, W. of Norseman, 1–17 Nov. 2003, C. Lambkin & J. Recsei, Eucalyptus woodland; ANIC bulk sample 2183 (ANIC). 1.1.1972, Australia-A.C.T., Canberra-Yarralumla Stirling Park, Coll. R. Kohout; 2 ex. (ANIC). S. Aust. Narrina Stn, 2 km SW Molkegna Bore, 30°57'22"S 138°45'12"E, 15–25 Mar. 1999, Flinders Ra surv PAT camp; openherbland/Grassland; No. 25-035902 (SAMA). 31°10'S 138°32'E, bet. Angorichina & Parachilna s.A., 23 Sept. 78 J.G.; crawling on the ground at night; No. 25-035931 (SAMA). Athelstone S.A., 24 Oct. 67, 7 pm–2 am UV light, J. Szent Ivany; No. 25-035976 (SAMA). K.J. Sandery, Highbury East, Sth. Aust., Nov 1967; No. 25-036021, 25-035975; 2 ex. (SAMA). S. Austr., Hilmfirth, Fulham..., Jan. 1908, JW Mellor; No. 25-036008 (SAMA). Sth. Aust.—Swan Reach, 1.7k WSW Kroehs Landg., 34°41'54"S 139°35'04"E, Pitfall 24–28 Aug. 2004, DEH, SWA 02402; Tussock Grassland-F/P, Phrabmites australis, Schoenopectus validus, Agrostis avenacea var.; No. 25-036047 (SAMA). Dalby Q., Mrs. F.H. Hobler; No. 25-035887, 035881; 2 ex. (SAMA). Beverley, W.A.; Ditto W. Australia; No. 25-035886 (SAMA). Bowen, Queensland, A. Simson; No. 25-035890 (SAMA). Australia: St. Albans, Vic. 11.V.58, B.P. Moore (ANIC). Auburn, NSW (ANIC).At light, Innamincka S.A., 18 Oct. 1962, J. Audley; No. 25-035932 (SAMA). W. Australia, Lancefield Gold Mines, Laverton, Dr. W.H. Ince, 1906-5 (BMNH). 39-920, Salmon Gums; Reg.no. 83262 (WAMP). R.P. McMillan, Spencers Brook, 20.7.48; Reg.no. 83264 (WAMP). 34-920, W. Midland; Reg.no. 83261 (WAMP). Norseman, W. Austr.; F.E. Wilson Collection; Col-65723 (NMV). NSW, Narrabri, Jan. 1990, leg. A. Smith, Hangay collection; Eunatalis lugubris Blackburn, 1890 , det. K. Markó, 2002 (HNHM). Cap York, Austral.; N. porcata F., det. J. Frivaldszky (HNHM). Australia, QLD, Dawson River/Kreuzung Capricorn HWY 11.11.1991, leg. R. Gerstmeier (RGCM). W-Australia, 8.11.1987, Station Creek, 127 km s Leinster, leg. R. Gerstmeier (RGCM). Mackay; Collect. Plason (RGCM). S.- Australien, Herrmannsburg a. Finkefluss, v. Leonhardi V.G. (ZMB). 85750; Hamb. S.-W. Austr. Exped. 1905, Stat. 95, Boorabbin, 3. VII.; 22; Natalis spinicornis Bl. (ZMB). N.W. of S.A., H. Basedow; Natalis spinicornis Blackb ; K 42987 , K 324405; 2 ex. (AM). B. Australia, A.H. Elston; A.H. Elston Collection; K 324407; Natalis spinicornis Bl., Id. by A.H. Elston, 550 (AM). Natalis . n.g, porcata, Spinl N. Holl. (NMPC). Natalis spinicollis Blackb., S.-W. Aust. (QM). Australien, N.S. Wales, Febr. 1903; Coll. C. le Deus; det. Corporaal 1946, Eunatalis spinicornis Blackb. (ZMUC). NS Wales, Heyne; Mus. Rothschild, 12-9- 1914; Spinicornis Black., N.S. Wales (ZMUC). Australia; Coll. D. Ach., 1.90; P. de Bosse (MSCN). Austral. mer., Victoria (ZMB). Australia, occid. 1192; Mus. Hauschild, 12-9-1914 (ZMUC). Australia, Old Coll.; K 324406, K 42987 ; 2 ex. (AM). N. Holl., (?); Natalis spinicornis Blackb. (ZMB). N. Vict.; Mus. Hauschild 12-9-1914 (ZMUC). Vict., Mus. Hauschild 12-9-1914 (ZMUC). Victoria; Museum Paris, (Coll. C.H. Schill), H. Donckier 1909 (MNHN). Australia mer., Coll. Kirsch (MTD). S. Australien, Gomolka. 75 467 (ZMB). Vict.; Natalis porcata F.; Andrewes, Bequest., B.M. 1922-221 (BMNH). Victoria, Barton; Museum Paris, Coll. Gorham, 1914; 3 ex. (MNHN). Queensland, Natalis spinicornis Blackb. ; Ex Museo E. Hintz; Museum Paris, ex Coll. R. Oberthür (MNHN). Nov: Holl:; Ex coll. Janson (MNHN). Australie; Museum Paris, Coll. M. Pic (MNHN). Victoria, Australie; Museum Paris. Coll. M. Pic (MNHN). Herrmannsburg, Central Australia, H.J. Nillier, 1911-311 (MNHN). Nat. Mus. Victoria, W. Australia; Natalis spinicornis Blackb., N.W. Australia; Nat. Mus. Victoria, C. French's Coll. 5.11.08; Col-65757, 65752; 2 ex. (NMV). S. Australia; Natalis spinicornis Blkb., Id. by A.H. Elston; F.E. Wilson Collection; Col-65756 (NMV). Australia, W. White; No. 25-035977 (SAMA). Guta, NSW (MZM). (?)Melton (MZM). 17802 (ZMB). Coolamundra; No. 0 35879 (SAMA). N.S. Wales; No. 25-035885; 2 ex. (SAMA). 42-456, Perth; Reg.no. 83263 (WAMP). Perth, April 1913, 7090; Reg.no. 83260 (WAMP). Sydney district, N.S.W., J.J.W. (HEC).

Diagnosis: E. spinicornis differs from all other species by the dentate elytral apices and a distinct longitudinal groove on the mesoventrite.

Length: (186 specimens measured) Total length: 20–31mm.

Head: Dark brown, with more or less short setae; with dense punctation and fine wrinkles; epistomal suture distinct; eyes very large and flattened; interocular space about one eye width; A11 acuminate.

Thorax: Pronotal length to width ratio 1.09:1; pronotum light brown to dark brown, with isolated pilosity; with weak punctation, sides wrinkled, dorsally partly with transverse wrinkles; grooves of sub-basal collar distinct, separated by a wide bar. Scutellum light brown to dark brown, with weak punctation.

Elytra: Elytral length to width ratio 2.71:1; elytra light brown to dark brown, basal half darker than apical half, with a light transverse fascia in the middle, with isolated pilosity; elytral base broader than pronotum, broader behind the middle, apex dentate; diameter of punctations in basal half double as wide as interstices, decreasing behind middle; punctations of tenth stria distinct up to shortly behind middle; punctations of ninth stria vestigial; elytral punctation with three nodules; interstices 4 and 6 conspicuously ridged behind the middle.

Lower surface: Light brown to dark brown, with dense pilosity and weak punctation; procoxal cavities opened posteriorly; metaventrite with an oblique row of weak wrinkles; mesoventrite with a distinct longitudinal groove; abdominal ventrites laterally with a light or yellowish impression.

Legs: Dark brown, with isolated pilosity, weak punctation and weak wrinkles; pro- and mesotibiae slightly bent.

Abdomen: Tegmen ventrally closed; phallic tip with a weak hook-like structure; spicular fork fused less than one-fifth.

Distribution: Mainly in the southern parts of Australia.

Seasonal occurrence: Collected throughout the year.

Figures

FIGURE 49. Eunatalis spinicornis: (A) Phallus lateral, (B) Phallus ventral, (C) Tegmen ventral, (D) Spicular fork, (E) Pygidium, (F) Ventrite VI.

FIGURE 50. Distribution of Eunatalis spinicornis.

FIGURE 58. (A) Eunatalis spadicea n. sp., (B) Eunatalis spinicornis, (C) Eunatalis titana, (D) Eunatalis villosa n. sp.