Edaphus parvipennis, Bernhauer, 1922

Herman, Lee H., 2001, Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the End of the Second Millennium IV. Staphylinine Group (Part 1) Euaesthetinae, Leptotyphlinae, Megalopsidiinae, Oxyporinae, Pseudopsinae, Solieriinae, Steninae, Bulletin of the American Museum of Natural History 2440 (265), pp. 1807-2440 : 1807-2440

parvipennis Bernhauer, 1922 , see: paraguayanus Bernhauer, 1923 .

parvulus L. Benick, 1914 , see: ageus Casey, 1884 .

pasticus L. Benick, 1938: 16 ( Stenus ; subgenus Hemistenus ; Type locality: G. Tanggamoes, de Giesting, Süd-Sumatra).

- DISTRIBUTION: Indonesia.

pastoralis Cameron, 1929b: 447 ( Stenus ; Type locality: Selangor: The Gap ).

— Scheerpeltz, 1933: 1163 ( Stenus ; subgenus Stenus ; catalog).

— Puthz, 1980a: 31 ( Stenus ; characters).

- DISTRIBUTION: Malaysia.

patens Casey, 1884 , see: pollens Casey, 1884 .

patruelis L. Benick, 1929 , see: submaculatus Bernhauer, 1915 .

pauliani Bernhauer, 1942 , see: matumbiensis Bernhauer, 1915 .

pauloensis Bernhauer, 1908d: 299 ( Stenus ; Type locality: Brasilien: S. Paulo).

— Bernhauer and Schubert, 1911: 176 ( Stenus ; subgenus Hypostenus ; catalog).

— Puthz, 1974a: 66, 69 ( Stenus ; characters; Brazil).

- DISTRIBUTION: Brazil.

paululus L. Benick, 1924: 251 ( Stenus ; Type locality: Tschita, Transbaikalien).

— Scheerpeltz, 1933: 1163 ( Stenus ; subgenus Stenus ; catalog).

— Puthz, 1966e: 298 ( Stenus ; synonym of innuptus ).

— Puthz, 1974b: 110 ( Stenus ; valid species; characters; Siberia).

— Campbell and Davies, 1991: 111 ( Stenus ; USA; Canada).

— Ryabukhin, 1999: 45 ( Stenus ; subgenus Stenus ; catalog; habitat; distribution; northeastern Russia).

- DISTRIBUTION: Russia, USA, Canada.

campbelli Puthz, 1973 h: 204 ( Stenus ; Type locality: USA: Alaska: Mi. 123 Taylor Highway ).

— Puthz, 1974a: 110 ( Stenus ; synonym of paululus ).

pauper Casey, 1884 , see: difficilis Casey, 1884 .

pauperculus Casey, 1884: 124 ( Stenus ; Type locality: locality uncertain but probably Michigan).

— Bernhauer and Schubert, 1911: 164 ( Stenus ; catalog).

— Puthz, 1971q: 244, 250, 254 ( Stenus ; characters; notes; Michigan; Massachusetts; Wisconsin). — Puthz, 1975g: 124 ( Stenus ; Quebec).

— Campbell and Davies, 1991: 111 ( Stenus ; Canada).

— Downie and Arnett, 1996: 492 ( Stenus ; characters; USA).

- DISTRIBUTION: USA, Canada.

brumalis Casey, 1884: 131 ( Stenus ; Type locality: Detroit, Michigan; Ann Arbor, Michigan). — Bernhauer and Schubert, 1911: 164 ( Stenus ; synonym of pauperculus ).

— Puthz, 1971q: 250 ( Stenus ; synonym of pauperculus ).

pauxillus Puthz, 1971: 227 ( Stenus ; subgenus Hypostenus ; Type locality: Congo: Kivu,

contr. S. Kahuzi, 2300 m).

- DISTRIBUTION: Congo.

payeri Krauss, 1899 , see: flavipes Stephens, 1833 .

peculiaripes Puthz, 1995e: 206 ( Stenus ; subgenus Tesnus ; Type locality: Brasilien: Para:

Rio Iriri Camp, 52°40′W, 3°50′S, ca 100 km S. Altamira).

- DISTRIBUTION: Brazil.

peculiaris Puthz, 1972 , see: phaeax Puthz, 1972 .

pedator Sharp, 1876: 367 ( Stenus ; Type locality: Rio Purus ).

— Bernhauer and Schubert, 1911: 184 (Stenus; subgenus Hemistenus; catalog).

— Hromádka, 1983: 121 ( Stenus ; characters).

- DISTRIBUTION: Brazil.

pedemontanus Puthz, 1994: 844 ( Stenus ; subgenus Parastenus ; Type locality: Italien: Piemont: Rima, im Val Sesia).

— Puthz and Zanetti, 1995: 18 ( Stenus ; Italy).

- DISTRIBUTION: Italy.

pekinensis Csiki, 1901 , see: alienus Sharp, 1874 .

pelidnus L. Benick, 1938a: 150 ( Stenus ; subgenus Hypostenus ; Type locality: Brasilien:

São Paulo).

- DISTRIBUTION: Brazil.

pendleburyi Cameron, 1950 , see: flavovittatus Champion, 1920 .

penelope Puthz, 1972 r: 498 ( Stenus ; subgenus Hypostenus ; Type locality: New Guinea NE: Simbai, Bismarck Range, 1700 m).

- DISTRIBUTION: New Guinea.

penicillifer Puthz, 2000: 208 ( Stenus ; subgenus Hypostenus ; Type locality: Ecuador: Pastaza Province: Puyo [1.5 km NE]).

- DISTRIBUTION: Ecuador.

peniculus L. Benick, 1938, see: basicornis Kraatz, 1859 .

peninsularis Cameron, 1929b: 451 ( Stenus ; subgenus Mesostenus ; Type locality: Selangor: The Gap. Penang).

— Scheerpeltz, 1933: 1195 ( Stenus ; subgenus Hemistenus ; catalog).

- DISTRIBUTION: Malaysia.

peraffinis Fauvel, 1895: 209 ( Stenus ; Type locality: Bírmanie, Carin Asciuii Ghécu , 1400–1500 m; [Note: See lectotype designation by Puthz, 1969]).

— Bernhauer and Schubert, 1911: 184 ( Stenus ; subgenus Hemistenus ; catalog).

— Cameron, 1930: 393 ( Stenus ; subgenus Mesostenus ; characters; Burma).

— Scheerpeltz, 1933: 1195 ( Stenus ; subgenus Hemistenus ; catalog).

— Puthz, 1969: 37 ( Stenus ; subgenus Parastenus ; lectotype designation: Carin Asciuii Ghecù, 1400–1500 m; characters).

— Puthz, 1981b: 152 ( Stenus ; characters).

- DISTRIBUTION: Myanmar.

perangustus Puthz, 1971 q: 252, 254 ( Stenus ; subgenus Nestus ; Type locality: California: Humboldt Co.: Fort Gaston, Hoopa Valley, Trinity River).

- DISTRIBUTION: USA.

peratus Cameron, 1930: 344 ( Stenus ; Type locality: Simla Hills. Chakrata district: Konain; Kanasar, alt. 7000 to 8000 feet).

— Scheerpeltz, 1933: 1163 ( Stenus ; subgenus Stenus ; catalog).

— Puthz, 1968j: 473 ( Stenus ; subgenus Parastenus ; characters; India).

— Abdullah and Qadri, 1968a: 304 ( Stenus ; characters; Pakistan).

— Puthz, 1976: 8 ( Stenus ; Nepal).

— Puthz, 1986k: 962 ( Stenus ; Pakistan).

- DISTRIBUTION: Pakistan, India, Nepal.

perbrevis L. Benick, 1949, see: lugens Casey, 1884 .

peregrinus Sharp, 1886: 656 ( Stenus ; Type locality: Panama, David in Chiriqui).

— Bernhauer and Schubert, 1911: 176 ( Stenus ; subgenus Hypostenus ; catalog).

— L. Benick, 1938c: 270 ( Stenus ; characters).

- DISTRIBUTION: Panama.

perexilis Notman, 1920a: 191 ( Stenus ; Type locality: Windsor, Broome Co., N.Y.).

— Scheerpeltz, 1933: 1163 ( Stenus ; subgenus Stenus ; catalog).

— Downie and Arnett, 1996: 491 ( Stenus ; characters; USA).

- DISTRIBUTION: USA.

perfidiosus Puthz, 1983: 11 ( Stenus ; subgenus Parastenus ; Type locality: Vietnam: Mt. Lang Bian, 1500–2000 m).

- DISTRIBUTION: Vietnam.

perforatus Casey, 1884: 169 ( Areus ; Type locality: Arizona, Morrison ).

— Bernhauer and Schubert, 1911: 176 ( Stenus ; subgenus Hypostenus ; catalog).

— L. Benick, 1938c: 277 ( Stenus ; characters).

- DISTRIBUTION: USA.

perhostilis Puthz, 2001: 12 ( Stenus ; subgenus Stenus ; Type locality: Costa Rica: Heredia:

27 km N Heredia, Rt. 9, 6000 feet. Also cited from Panama and Guatemala).

- DISTRIBUTION: Guatemala, Costa Rica, Panama.

perillustris Puthz, 1972: 26 ( Stenus ; Type locality: Madagaskar: Mt. d’Ambre ).

— Puthz, 1992a: 14 ( Stenus ; Madagascar).

- DISTRIBUTION: Madagascar.

peringueyi L. Benick, 1933: 308 ( Stenus ; Type locality: Südafrika: Kapkolonie: Dumbrody ). — Puthz, 1971: 137 ( Stenus ; characters; South Africa; Angola).

— Puthz, 1971k: 137 ( Stenus ; South Africa).

— Hammond, 1975a: 9 ( Stenus ; notes; South West Africa; Botswana).

- DISTRIBUTION: South Africa, Botswana, Namibia, Angola, Congo, Tanzania, Cameroon.

fossicollis L. Benick, 1951: 179 ( Stenus ; Type locality: Ost-Afrika: Kondoa, 1300 m).

— Puthz, 1971: 139 ( Stenus ; subspecies of peringueyi ; characters; Tanzania).

kamerunicola Puthz, 1967 r: 1009 ( Stenus ; subgenus Stenus ; subspecies of fossicollis ; Type locality: Cameroun: marais du Noung).

— Puthz, 1971: 140 ( Stenus ; subspecies of peringueyi ; Cameroon).

dubitabilis Puthz, 1971: 141 ( Stenus ; subspecies of peringueyi ; Type locality: Tanzania: Kilimandjaro, versant Sud-Est, zone inférieure, New-Moschi, 800 m).

kolweziensis Puthz, 1971: 141 ( Stenus ; subspecies of peringueyi ; Type locality: Congo: Katanga, Kolwezi, Manika, Dembo desséché).

peripherus Korge, 1971 , see: melanarius Stephens, 1833 .

periscelidifer Puthz, 1984e: 568 ( Stenus ; subgenus Hypostenus ; Type locality: Kerala: Anaimalai Hills, 18 km au nord de Valparai, 1250 m).

- DISTRIBUTION: India.

permixtus Fagel, 1967a: 355 ( Stenus ; subgenus Parastenus ; Type locality: Caucas. occ. Circassien).

— Ryvkin, 1990: 187 ( Stenus ; notes; Russia).

- DISTRIBUTION: Russia.

pernix L. Benick, 1929: 261 ( Stenus ; subgenus Hypostenus ; Type locality: Montalban, Luzon). — Scheerpeltz, 1933: 1180 ( Stenus ; subgenus Hypostenus ; catalog).

- DISTRIBUTION: Philippines.

pernobilis Puthz, 1998c: 240 ( Stenus ; subgenus Parastenus ; Type locality: Sulawesi: Gunang Mogogonia Pa, 1080 m, Utara).

- DISTRIBUTION: Indonesia.

peroculatus Puthz, 1986 , see: comma LeConte, 1863 .

perplexus Puthz, 1973d: 73 ( Stenus ; subgenus Tesnus ; Type locality: Borneo: W. Sarawak:

Mt. Matang).

— Puthz, 1988: 675 ( Stenus ; Borneo).

- DISTRIBUTION: Borneo.

perpulcher Solsky, 1875: 24 ( Stenus ; Type locality: Pérou, Maraquia ).

— Bernhauer and Schubert, 1911: 164 ( Stenus ; catalog).

— Puthz, 2000: 169 ( Stenus ; subgenus Stenus ; characters; Ecuador; Bolivia).

- DISTRIBUTION: Bolivia, Peru, Ecuador.

perroti Puthz, 1981b: 157 ( Stenus ; subgenus Parastenus ; Type locality: Vietnam [ Tonkin]: Tam Do. Also cited from Burma).

— Rougemont, 1981a: 342 ( Stenus ; characters of male; Burma).

— Puthz, 1981a: 18 ( Stenus ; Vietnam).

— Rougemont, 1983c: 45 ( Stenus ; Thailand).

— Puthz, 1983: 7 ( Stenus ; Vietnam; China).

- DISTRIBUTION: Vietnam, Myanmar, China.

perrotianus Puthz, 1998: 386 ( Stenus ; subgenus Parastenus ; Type locality: Vietnam: Tam Dao). - DISTRIBUTION: Vietnam.

perrugosus L. Benick, 1917b: 297 ( Stenus ; subgenus Stenus ; Type locality: Columbien:

El Tambo, 6000′, Bogota, Terra templ.).

— Scheerpeltz, 1933: 1163 ( Stenus ; subgenus Stenus ; catalog).

— L. Benick, 1938c: 262 ( Stenus ; characters).

— Puthz, 1969j: 57 ( Stenus ; aedeagus).

- DISTRIBUTION: Colombia.

persicus Puthz, 1981 i: 705 ( Stenus ; subgenus Parastenus ; Type locality: Iran: Kermanshah: Kenesht/ Kermanshah, 34°29′N 47°09′E).

— Ryvkin, 1990: 212 ( Stenus ; notes; Iran).

- DISTRIBUTION: Iran.

persimilis Cameron, 1930: 367 ( Stenus ; subgenus Hypostenus ; Type locality: Burma:

Ruby Mines).

— Rougemont, 1987: 707 ( Stenus ; subgenus Hypostenus ; Thailand).

- DISTRIBUTION: Myanmar, Thailand.

personatus L. Benick, 1928d: 52 ( Stenus ; Type locality: Nord-Amerika, Black Mts.,

Mount Mitchell, N.C.).

— Scheerpeltz, 1933: 1195 ( Stenus ; subgenus Hemistenus ; catalog).

- DISTRIBUTION: USA.

perssoni Puthz, 1968 i: 456 ( Stenus ; subgenus Parastenus ; Type locality: Brasilien:

Sta. Catharina: Nova Teutonia).

- DISTRIBUTION: Brazil.

pertinax Casey, 1884: 195 ( Areus ; Type locality: Wyoming Territory ).

— Bernhauer and Schubert, 1911: 176 ( Stenus ; subgenus Hypostenus ; catalog).

— Puthz, 1974d: 166 ( Stenus ; notes).

- DISTRIBUTION: USA.

lucidus Casey, 1884: 196 ( Areus ; Type locality: Oak Grove , California).

— Fall, 1901: 71 ( Stenus ; California).

— Bernhauer and Schubert, 1911: 175 ( Stenus ; synonym of leviceps ).

— Puthz, 1974d: 166 ( Stenus ; synonym of pertinax ).

pertusus Fairmaire and Germain, 1861: 444 ( Stenus ; Type locality: Chiloe; [Note: See lectotype designation by Puthz, 1973g]).

— Fauvel, 1867a: 19 [= 1868: 24] ( Stenus ; characters; Chile).

— Bernhauer and Schubert, 1911: 176 ( Stenus ; subgenus Hypostenus ; catalog).

— Coiffait and Saiz, 1968: 400 ( Stenus ; subgenus Hypostenus ; characters; Chile).

— Puthz, 1972g: 128 ( Stenus ; subgenus Hypostenus ; characters; Chile).

— Puthz, 1973g: 177 ( Stenus ; lectotype designation; notes).

- DISTRIBUTION: Chile.

peruvianus Bernhauer, 1908d: 299 ( Stenus ; Type locality: Peru: Puno).

— Bernhauer and Schubert, 1911: 176 ( Stenus ; subgenus Hypostenus ; catalog).

— Puthz, 1974a: 66 ( Stenus ; characters).

- DISTRIBUTION: Peru.

petrii L. Benick, 1913 , see: phyllobates Penecke, 1901 .

pettiti Casey, 1884, see: angustus Casey, 1884.

pfeifferi Puthz, 1998c: 237 ( Stenus ; subgenus Hypostenus ; Type locality: Borneo: Sarawak: Umg Marudi, 4°10′N, 114°20′E, Ufer des Baram. Also cited from Brunei).

- DISTRIBUTION: Borneo.

pflegeri R. Dvořák and Havelka, 1951: 4 ( Stenus ; subgenus Parastenus ; Type locality: Caucas. occ.: Circasien).

— Fagel, 1967a: 352 ( Stenus ; subgenus Parastenus ; characters; Caucasus).

— Puthz, 1983e: 359 ( Stenus ; Caucasus).

— Ryvkin, 1990: 184 ( Stenus ; notes; Russia).

- DISTRIBUTION: Russia.

phaeax Puthz, 1972 r: 504 ( Stenus ; subgenus Hypostenus ; Type locality: New Guinea: Morobe: Herzog Mts., Vagau, ca 4000 ft.).

- DISTRIBUTION: New Guinea.

peculiaris Puthz, 1972 r: 505 ( Stenus ; subgenus Hypostenus ; subspecies of phaeax ; Type locality: Wau Creek, 1200–1300 m).

phaenomenalis Bernhauer, 1919: 362 ( Stenus ; Type locality: Luzon: Los Banos,

Mt. Makiling).

— L. Benick, 1929: 246 ( Stenus ; subgenus Hypostenus ; characters; Philippines).

— Scheerpeltz, 1933: 1180 ( Stenus ; subgenus Hypostenus ; catalog).

- DISTRIBUTION: Philippines.

orousseti Puthz, 1984c: 171 ( Stenus ; subgenus Hypostenus ; subspecies of phaenomenalis ; Type locality: Philippinen: Mindoro: Puerto Galera, 300 m).

phaeopus Stephens, 1833 , see: bifoveolatus Gyllenhal, 1827 .

philippinus Bernhauer, 1912 g: 249 ( Stenus ; Type locality: Not cited but presumably

The Philippines).

— L. Benick, 1929: 263 ( Stenus ; subgenus Hypostenus ; characters; Philippines).

— Scheerpeltz, 1933: 1180 ( Stenus ; subgenus Hypostenus ; catalog).

- DISTRIBUTION: Philippines.

phyllobates Penecke, 1901: 14 ( Stenus ; subgenus Nestus ; Type locality: Lobnitzbach in

der Nähe der “Alten Glashütten; Bachergebirge in Steiermark; Rodna-Gebirge

in Siebenbürgen).

— Bernhauer and Schubert, 1911: 164 ( Stenus ; catalog).

— L. Benick, 1929a: 40 ( Stenus ; subgenus Nestus ; cited as phyllobatus; characters in key).

— Scheerpeltz, 1933: 1163 ( Stenus ; subgenus Stenus ; catalog).

— Szujecki, 1961: 30 ( Stenus ; subgenus Nestus ; characters; Poland).

— Horion, 1963: 357 ( Stenus ; subgenus Nestus ; Romania; Austria).

— Lohse, 1964: 116 ( Stenus ; subgenus Nestus ; characters; central Europe).

— Smetana, 1964a: 50 ( Stenus ; subgenus Nestus ; characters).

— Puthz, 1966f: 119 ( Stenus ; subgenus Nestus ; notes).

— Puthz, 1968e: 98 ( Stenus ; subgenus Nestus ; notes).

— Szujecki, 1969a: 252 ( Stenus ; notes; Poland).

— Puthz, 1971l: 170 ( Stenus ; distribution of subspecies).

— Puthz, 1971v: 92 ( Stenus ; notes).

— Puthz, 1976j: 190 ( Stenus ; distribution in Germany).

— Puthz, 1982e: 85 ( Stenus ; distribution of polytypic species).

— Lucht, 1987: 97 ( Stenus ; checklist; Germany; Poland; Czechoslovakia; Austria).

— Lohse and Lucht, 1989: 143 ( Stenus ; notes).

— Dauphin, 1993: 179, 186 ( Stenus ; subgenus Nestus ; characters).

— Hromádka, 1993: 44 ( Stenus ; subgenus Nestus ; Czech Republic; Slovakia).

— Puthz and Zanetti, 1995: 18 ( Stenus ; Italy).

— Jelínek, 1999: 157 ( Stenus ; subgenus Nestus ; collecting note; Czech Republic).

- DISTRIBUTION: Europe.