Ogcodes kunkunche Barahona-Segovia, 2020

Barahona-Segovia, Rodrigo M., Guzmán, Vicente Valdés, Barceló, Matías & Pañinao-Monsálvez, Laura, 2020, A new spider fly (Diptera: Acroceridae: Ogcodinae: Ogcodes Latreille) from Chiloé Island’s evergreen forest and new distributional records for other spider flies in Chile, Zootaxa 4779 (1), pp. 51-64 : 54-55

Ogcodes kunkunche Barahona-Segovia sp. nov.

Figs. 1–2 , 5–8

Holotype. [Holotype without mid and hind legs. Deposited in MNHNCL, with the following labels]: (1) “Caulin, Chacao, Chiloé / 14.ii.2019 / Leg. Marco Prado” and (2) “ Holotypus / Ogcodes kunkunche / sp. nov. ♂ / det. Bara- hona-Segovia 2019” (red).

Diagnosis: Scutum shiny black with scutellum golden-yellow. Mid legs missing as well as hind tibiae and tarsi (see notes below). Fore femur bicolor with anterior part black and swollen throughout, while posterior femur yellow and not swollen. Tibiae and tarsi 1–4 golden yellow with fifth tarsus blackish. Tergite 1 with thin and pale white fascia in the posterior margin and tergites 2–6 golden-yellow with fascia of irregular shape near the lateral margins.

Description: Total length: 4.1 mm; head height and width: 1.1 and 0.8 mm, respectively; thorax height and width: 1.8 mm and 1.7, mm respectively; Wing length: 3.2 mm. MALE. Head: Spheroidal; eyes dull black lacking ommatrichia; vertex large and black with black ocelli located anterolaterally; ocellar triangle with two darkish, scarcely protuberant ocelli occupying 50% of vertex; clypeus black; ( Fig. 2 ); occipital setae black; antenna black, flagellum with four black setae ( Fig. 3 ); Thorax: scutum shiny black, with large erected black pilosity continuous and sparsely distributed throughout ( Figs. 5–6 ); anepisternum, anepimeron, katepisterum, metasternum, meron and katatergite shiny black without hairs or with very scarce, short and black pilosity ( Fig. 5 ); scutellum golden-yellow, with erected hairs of the same color ( Figs. 5–6 ); Legs: mid legs missing as well as hind tibiae and tarsi due to manipulation of collector (see notes below); coxa and trochanter shiny black ( Fig. 5 ); fore femora swollen with dense pilosity, with proximal area blackish and a golden-yellow sickle-shape marking; hind femora enlarged, not swollen, with few golden-yellow maculae distally; large and black hairs present ( Fig. 5 ); hind tibiae golden-yellow and thickened posteriorly, with short, yellow hairs distributed throughout; hind tarsi with golden-yellow color, excepting the fifth tarsus, which is darkish brown; claws darkish brown color ( Fig. 5 ); Wing: with hyaline appearance, veins brown in color; cells c and sc with light brown color; veins M 1 and M 2 absent; M 4 and CuA present, light brown; crossvein m–cu absent; halter golden-yellow; calypter rim shiny black and interior part is brownish ( Fig. 7 ). Abdomen: Triangular ( Fig. 6 ); tergite 1 completely shiny black with posterior thin white-yellow pale fascia; tergites 2–6 shiny black with posterior golden-yellow fasciae; the first three fasciae cover about one third of each segment, thickened toward the abdominal margins; fascia of tergite 4 more enlarged than those of tergites 1 and 2; golden-yellow fascia covering approximately 50% and 80% of tergites 5 and 6, respectively ( Fig. 6 ); Abdominal fascia on tergites 2–5 not reaching the anterior margin; sclerites dull black with whitish fascia in the posterior margin; Genitalia: partially everted (not dissected for this study); distal portion of aedeagus and gonopore rounded with a white mark; apex of the clasper brown ( Fig. 8 ).

Female: Unknown.

Etymology: The specific epithet ‘ kunkunche ’ refers to the southern Mapuche people named ‘Cuncos’. The “cuncos” or “puncos” (in mapudungun language = kunkunche ) were a sedentary indigenous people, frequently considered to be the southern-most branch of the Mapuche people. They lived in southern Chile from the Maullín River to the northern part of Chiloé and the neighboring islands. This indigenous people developed agriculture, cultivating several crops including potatoes and corn. Also, they were dedicated to recollecting marine mollusks and hunting marine fishes, sea lions and other marine mammals. The name “ kunkunche ” was chosen for this spider fly species because this indigenous people were the group most hostile to the Spanish colonizers, and this represents the persistence of the new species described herein in the fragmented landscape that currently dominates the main Chiloé Island.

Distribution: Caulín, 41.816013° S, 73.596358° W, 48 m, Chacao, main Chiloé Island ( Fig. 1 ).

Remarks: Currently, Caulín is a rural town located in northern Chiloé Island with fragmented native forest cleared for livestock, grassland or crop use. The remaining forest is mainly secondary growth, composed primarily by Myrceugenia spp., Luma apiculata and Eucryphia cordifolia . The holotype specimen was collected on flowers of an isolated E. cordifolia tree at mid-day in semi-clouded weather.

Notes: the new species has damage in the legs produced by bad manipulation of collector. Hind legs are complete, but mid legs are totally missing. In addition, fore legs present both femora, but tibiae and tarsi are absent.

Figures

FIGURE 1. Distribution maps of the five Chilean species of Ogcodes.

FIGURES 2–4. Ogcodes kunkunche Barahona-Segovia sp. nov.: (2) male, head, anterior view; (3) male, antenna; (4) Ogcodes porteri Schlinger, male, head, anterior view.

FIGURES 5–6. Ogcodes kunkunche Barahona-Segovia sp. nov.: (2) male, lateral view; (3) male, dorsal view.

FIGURES 7–8. Ogcodes kunkunche Barahona-Segovia sp. nov.: (6) male, left, wing venation; (7) male, genitalia, lateral view.