Oropycnodus ponsorti ( Heckel, 1854 ) Poyato-Ariza F. J. & Wenz S., 2002

Poyato-Ariza F. J. & Wenz S., 2002, A new insight into pycnodontiform fishes, Geodiversitas 24 (1), pp. 139-248 : 223

Oropycnodus ponsorti ( Heckel, 1854) n. comb.

( Figs 4B ; 11A, B ; 17A, B ; 29A, B ; 42A, B ; 46B ) LECTOTYPE. — NMW 1854 /XXXIX/38 ( Heckel 1856: pl. 11, fig. 1; Figs 4B ; 17A, B ; 42A, B ).

PARALECTOTYPES. — NMW 1854/XXXIX/39 ( Heckel 1856; Fig. 46B ) and 1854/XXXIX/40 ( Heckel 1856: pl. 11, fig. 2; Fig. 11A, B ).

REFERRED SPECIMENS. — MNHN MTA 3, 4, 5, 6, 7, 9, 10, 11, 12, 13, 15, 16, 17, 38, 39, 40, 41, 42 ( Fig. 29A ), 43, 44, 45, 46, 47, 48, 49; NHML 30035, 30036, 30037, 30038, 30039, 30040, 30042, 30043, 30044, 30045, 30046, 30047, P1638.

TYPE HORIZON. — Paleocene (Montian).

TYPE LOCALITY. — Mont Aimé, Chalons-sur-Marne, France.

REVISED DIAGNOSIS. — As for genus (monotypic genus).

Figures

FIG. 4. — Forms previously referred to the genus Palaeobalistum Blainville, 1818; A, P. orbiculatum Blainville, 1818, type species, holotype, MNHN BOL 0523. Photo Serrette; B, Oropycnodus ponsorti n. comb., lectotype, NMW.1854.XXXIX.38. Photo Schumacher, courtesy Ortwin Schultz; C, D, Abdobalistum thyrsus n. gen., n. sp., part and counterpart of the holotype, NMHL P.9830a-b. Photos courtesy P. Forey. Scale bars: A, 5 cm; B-D, 2 cm.

FIG. 11. — Oropycnodus ponsorti n. comb., paralectotype, NMW 1854.XXXIX.40, left side, lateral view; A, complete specimen. Photo Schumacher, courtesy Ortwin Schultz; B, detail of the posterior region of the skull roof. Abbreviations: dmfe, dermocranial fenestra; drs, dorsal ridge scales; drs1, dorsal ridge scale 1; end, endocranium (posteriorly exposed); Fr, frontal; papr, parietal process; pcla postcephalic lacuna; Socc, dermal supraoccipital. Scale bars: A, 1 cm; B, 5 mm.

FIG. 17. — Skull of Oropycnodus ponsorti n. comb., lectotype, NMW 1854.XXXIX.38, right side, lateral view; A, complete skull. Photo Schumacher, courtesy Ortwin Schultz; B, camera lucida drawing showing detail of the opercular region. Abbreviations: Br, branchiostegal rays; Cl, cleithrum; Dhy, dermohyomandibular; Op, operculum; pcf, pectoral fin; Pop, preoperculum; Scl, supracleithrum. Scale bars: A, 1 cm; B, 2 mm.

FIG. 29. — Oropycnodus ponsorti n. comb.; A, caudal region of specimen MNHN MTA 42. Photo Serrette; B, restoration of the caudal skeleton, mostly from camera lucida drawings of specimen in A and of lectotype NMW 1854/XXXIX/38 and paralectotype NMW 1854/XXXIX/40. Arrows on B indicate upper and lower principal caudal fin rays. Both left side. Abbreviations: aax, anal axonost; cfr, caudal fin rays; epel, epichordal elements; ha, haemal arcocentrum; hs, haemal spine; hyel, hypochordal elements; na, neural arcocentrum; ns, neural spine; sf, sagittal flange; ud, urodermal. Scale bars: A, 5 mm; B, 2 mm.

FIG. 42. — Oropycnodus ponsorti n. comb., non-mosaic cloaca with bifid scale; A, cloacal region of the lectotype, NMW 1854.XXXIX.38. Photo Schumacher, courtesy Ortwin Schultz; B, restoration of the cloacal region, mostly after a camera lucida drawing of the lectotype, illustrated in A, with information taken from camera lucida drawings of the paralectotypes, NMW 1854.XXXIX.39 and NMW 1854.XXXIX.40, and, topotypes NHML P1638, P3004, and P.30037. Both right side, lateral view. Abbreviations: aax, anal axonosts; af, anal fin; bcls, bifid cloacal scale; co, comma-shaped cloacal scales; csc, complete scales; cv, cloacal vestibule; opv, opening for pelvic fin; pcb, post-coelomic bone; pcn, post-cloacal notch; r, ribs; rsc, reduced scales; vks, ventral keel scales. Scale bars: A, 2 mm; B, 5 mm.

FIG. 46. — Camera lucida drawings of branchial teeth as seen in A, the gill chamber of specimen MNHN HAK 298 of Ichthyoceros spinosus Gayet, 1984; B, the gill chamber of paralectotype NMW 1854/XXXIX/39 of Oropycnodus ponsorti n. comb.; C, disarticulated from specimen FSL 400047 of Proscinetes sp. (C1, in gill chamber; C2, outside the body, anteroventral to left dentary; C3, same position, closer to left dentary). Scale bars: 1 mm.